Liver (color)

Last updated
Liver
 
Gtk-dialog-info.svg    Color coordinates
Hex triplet #674C47
sRGB B (r, g, b)(103, 76, 71)
HSV (h, s, v)(9°, 31%, 40%)
CIELChuv (L, C, h)(35, 19, 22°)
Source ISCC-NBS
ISCC–NBS descriptor Grayish reddish brown
B: Normalized to [0–255] (byte)

At right is displayed the color traditionally called liver.

Contents

The first recorded use of liver as a color name in English was in 1686. [1]

Liver may also refer to a group of certain types of dark brown color in dogs and horses. Said nomenclature may also refer to the color of the organ.

In dogs

A Labrador Retriever. Note the brown nose, and the yellow in the eyes. In Labradors this color is called "Chocolate," but it is the same as "Liver." Labrador Retriever chocolate Hershey sit (cropped).jpg
A Labrador Retriever. Note the brown nose, and the yellow in the eyes. In Labradors this color is called "Chocolate," but it is the same as "Liver."
Liver (Dogs)
 
Gtk-dialog-info.svg    Color coordinates
Hex triplet #5D3B1A
sRGB B (r, g, b)(93, 59, 26)
HSV (h, s, v)(30°, 72%, 36%)
CIELChuv (L, C, h)(28, 33, 41°)
Source[Unsourced]
ISCC–NBS descriptor Dark brown
B: Normalized to [0–255] (byte)

In dogs, the Liver color is caused by dilution of the eumelanin (black) pigment by the B locus. The gene that causes Liver is recessive, so a BB or Bb dog has normal black pigment. Only a bb dog is Liver. There are several different recessive b genes, but they all turn the coat brown. They are only distinguishable through genetic testing. [2] Liver may also be called different names such as Brown, Chocolate, or Red. Red is a very misleading term, and should be avoided when referring to the color liver because it can be confused for phaeomelanin pigment. This second pigment is what colors all the "true red" or yellow parts of a dog.

Can Setter dog GFDL.jpg
An Irish Setter. This dog is not a liver, but a recessive red with very dark pigment.
Rough Collie 600.jpg
A Rough Collie. This dog is also not a liver, although it may be mistaken for it. It is a Black and Red/Yellow dog with the shaded sable pattern.
Neither of these dogs is a Liver.

Liver dilution can also be combined with other genes that dilute a dog's coat. The D locus dilutes black pigment to a blue-grey color usually called Blue. It is a recessive trait like Liver. When a dog has both Liver and Blue dilution, it will appear to be a light, warm-gray color known as Isabella. This is the color of Weimaraners. This color also has alternate names such as Liliac or Silver. When Liver is combined with Merle, the eumelanin (black) pigment is further diluted in random patches. This usually creates a light reddish-gray dog with dark brown patches. This color is often called "Red Merle" even though that is misleading. A more correct name would be Liver Merle or Chocolate Merle.

A German Wirehaired Pointer. This dog is a liver. It is also a piebald with roaning. GermanWirehrPtr1 wb.jpg
A German Wirehaired Pointer. This dog is a liver. It is also a piebald with roaning.

There are many other dog colors that can look like brown, but are not Liver. A dog that is very dark red, like an Irish Setter may sometimes appear brown. Dogs with a blend of Black and Red/Yellow fur can also look like they have brown areas. An easy way to tell if a dog is Liver or not is to look at their nose. Eumelanin (black) pigment colors a dog's nose, so a Liver dog will have a Liver colored nose. If the nose is black, the dog is not a liver. A pink nose has nothing to do with liver dilution, and will not help determine if a dog is liver or not. Liver will also dilute a dog's brown eyes to amber/yellow.

It is also possible for a dog to be a Liver, and not appear brown. A dog that is recessive red cannot produce eumelanin (black) pigment in their fur. Since Liver is a dilution of black pigment, a recessive red Liver dog will appear to be a shade of Red, Yellow, or Cream depending on the intensity of the dog's phaeomelanin (red) pigment. A recessive red Liver will still be born with a Liver nose and amber/yellow eyes. It is common for the nose of any recessive red dog to fade to pink as they grow older. Livers, like any dog, can also be covered with varying amounts of white which will hide the color of the fur.

Liver chestnut in horses

An example of a dark liver chestnut-colored horse. Tori horse universal.jpg
An example of a dark liver chestnut-colored horse.
Liver Chestnut (horses)
 
Gtk-dialog-info.svg    Color coordinates
Hex triplet #543D37
sRGB B (r, g, b)(84, 61, 55)
HSV (h, s, v)(12°, 35%, 33%)
CIELChuv (L, C, h)(28, 16, 26°)
Source[Unsourced]
ISCC–NBS descriptor Dark grayish brown
B: Normalized to [0–255] (byte)

In horses, liver chestnut is a chocolate-colored chestnut horse. A dark liver chestnut has the same recessive base genetics as a regular chestnut, but the shade is a dark brown rather than the reddish or rust color more typical of chestnut. A horse that appears to be a dark liver chestnut but has a flaxen-colored mane and tail, sometimes colloquially though incorrectly called a "chocolate palomino", could be genetically chestnut but could also be a black horse manifesting the silver dapple gene. Silver dapple is a dilution gene that acts in a manner similar to the liver dilution in dogs.

The darkest liver chestnuts may be confused for black, but such horses are distinguished from other phenotypes by the absence of black hairs, and obviously brown or reddish legs, muzzles, flanks and other "soft" areas. The genotypes that determine specific varieties of chestnut are not known, but the genetics that darken the coat may have a recessive mode of inheritance. The sooty gene may also be one factor in creating the darker color, though sooty coloration is generally not evenly distributed throughout the coat. Genetic testing can distinguish a chestnut from a black or a bay horse. [3]

Other dilution genes in horses analogous to those creating dilution shades in dogs do not create a liver color in horses. These include the dun gene, which produces a tan-colored or "blue"/gray solid colored coat and the champagne gene which tends to lighten the coat more than the mane or tail.

Liver (organ)

An image of a healthy sheep's Liver; human livers are much the same color Leber Schaf.jpg
An image of a healthy sheep's Liver; human livers are much the same color
Liver (organ)
 
Gtk-dialog-info.svg    Color coordinates
Hex triplet #6C2E1F
sRGB B (r, g, b)(108, 46, 31)
HSV (h, s, v)(12°, 71%, 42%)
CIELChuv (L, C, h)(27, 47, 20°)
Source[Unsourced]
ISCC–NBS descriptor Deep reddish brown
B: Normalized to [0–255] (byte)

This is the color of a healthy human liver. It may range from brown to reddish brown, and the color represented in the adjacent box is the gross average of these shades. A yellowish or greenish liver may indicate jaundice or a similar condition, a dark brown color may indicate alcohol poisoning,[ citation needed ] a black color can be seen in livers affected by Dubin–Johnson syndrome, [4] and white or grayish tones may indicate cancer.[ citation needed ] It is unknown why the color of dogs and horses came to be known by the term "Liver", as these tones indicate an unhealthy liver.[ citation needed ]

These healthy tones usually indicate blood flow, which is why livers and other meat turn grayish brown when cooked. [5]

Dark liver (web)

Liver (web) (#534B4F)
#534B4F

At right is displayed the color dark liver (web).

This is the shade of dark liver that is the unofficial web color called liver that is traditionally used in web site design.

In nature

See also

Related Research Articles

<span class="mw-page-title-main">Palomino</span> Genetic color in horses

Palomino is a genetic color in horses, consisting of a gold coat and white mane and tail; the degree of whiteness can vary from bright white to yellow. The palomino color derived from the inter-breeding of Spanish horses with those from the United States. Genetically, the palomino color is created by a single allele of a dilution gene called the cream gene working on a "red" (chestnut) base coat. Palomino is created by a genetic mechanism of incomplete dominance, hence it is not considered true-breeding. However, most color breed registries that record palomino horses were founded before equine coat color genetics were understood as well as they are today, therefore the standard definition of a palomino is based on the visible coat color, not heritability nor the underlying presence of the dilution gene.

<span class="mw-page-title-main">Dilution gene</span> Gene that lightens the coat colour of certain animals

A dilution gene is any one of a number of genes that act to create a lighter coat color in living creatures. There are many examples of such genes:

<span class="mw-page-title-main">Cat coat genetics</span> Genetics responsible for the appearance of a cats fur

Cat coat genetics determine the coloration, pattern, length, and texture of feline fur. The variations among cat coats are physical properties and should not be confused with cat breeds. A cat may display the coat of a certain breed without actually being that breed. For example, a Neva Masquerade could wear point coloration, the stereotypical coat of a Siamese.

<span class="mw-page-title-main">Bay (horse)</span> Hair coat color of horses

Bay is a hair coat color of horses, characterized by a reddish-brown or brown body color with a black point coloration on the mane, tail, ear edges, and lower legs. Bay is one of the most common coat colors in many horse breeds.

<span class="mw-page-title-main">Brindle</span> Coat coloring pattern in some animals

Brindle is a coat coloring pattern in animals, particularly dogs, cattle, guinea pigs, cats, and, rarely, horses. It is sometimes described as "tiger-striped", although the brindle pattern is more subtle than that of a tiger's coat.

<span class="mw-page-title-main">Point coloration</span> Coloration of animal coat/fur

Point coloration is animal coat coloration with a pale body and relatively darker extremities, i.e. the face, ears, feet, tail, and scrotum. It is most recognized as the coloration of Siamese and related breeds of cat, but can be found in dogs, rabbits, rats, sheep, guinea pigs and horses as well.

Canine terminology in this article refers only to dog terminology, specialized terms describing the characteristics of various external parts of the domestic dog, as well as terms for structure, movement, and temperament. This terminology is not typically used for any of the wild species or subspecies of wild wolves, foxes, coyotes, dholes, jackals or the basal caninae. Dog terminology is often specific to each breed or type of dog. Breed standards use this terminology in the description of the ideal external appearance of each breed, although similar characteristics may be described with different terms in different breeds.

<span class="mw-page-title-main">Cream gene</span> Gene for several horse coat colors

The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, if they are homozygous. Similarly, horses with a bay base coat and the cream gene will be buckskin or perlino. A black base coat with the cream gene becomes the not-always-recognized smoky black or a smoky cream. Cream horses, even those with blue eyes, are not white horses. Dilution coloring is also not related to any of the white spotting patterns.

<span class="mw-page-title-main">Champagne gene</span> Simple dominant allele responsible for a number of rare horse coat colors

The champagne gene is a simple dominant allele responsible for a number of rare horse coat colors. The most distinctive traits of horses with the champagne gene are the hazel eyes and pinkish, freckled skin, which are bright blue and bright pink at birth, respectively. The coat color is also affected: any hairs that would have been red are gold, and any hairs that would have been black are chocolate brown. If a horse inherits the champagne gene from either or both parents, a coat that would otherwise be chestnut is instead gold champagne, with bay corresponding to amber champagne, seal brown to sable champagne, and black to classic champagne. A horse must have at least one champagne parent to inherit the champagne gene, for which there is now a DNA test.

<span class="mw-page-title-main">Silver dapple gene</span>

The silver or silver dapple (Z) gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color, and a black body to a chocolaty brown, sometimes with dapples. It is responsible for a group of coat colors in horses called "silver dapple" in the west, or "taffy" in Australia. The most common colors in this category are black silver and bay silver, referring to the respective underlying coat color.

<span class="mw-page-title-main">Equine coat color genetics</span> Genetics behind the equine coat color

Equine coat color genetics determine a horse's coat color. Many colors are possible, but all variations are produced by changes in only a few genes. Bay is the most common color of horse, followed by black and chestnut. A change at the agouti locus is capable of turning bay to black, while a mutation at the extension locus can turn bay or black to chestnut.

<span class="mw-page-title-main">Chestnut (horse color)</span> Horse coat color

Chestnut is a hair coat color of horses consisting of a reddish-to-brown coat with a mane and tail the same or lighter in color than the coat. Chestnut is characterized by the absolute absence of true black hairs. It is one of the most common horse coat colors, seen in almost every breed of horse.

<span class="mw-page-title-main">Equine coat color</span> Horse coat colors and markings

Horses exhibit a diverse array of coat colors and distinctive markings. A specialized vocabulary has evolved to describe them.

<span class="mw-page-title-main">Black horse</span> Horse coat color

Black is a hair coat color of horses in which the entire hair coat is black. Black is a relatively uncommon coat color, and it is not uncommon to mistake dark chestnuts or bays for black.

<span class="mw-page-title-main">Smoky black</span>

Smoky black or black carrying cream is a coat color of horses which has the same phenotype as black. Smoky black is produced by the action of a heterozygous cream gene on an underlying black coat color. Therefore, smoky black is a member of the cream family of coat color dilutions, and found in horse populations that have other cream-based colors such as palomino, buckskin, perlino, cremello and smoky cream. All smoky blacks must have at least one parent with the cream gene, and a smoky black can only be verified through DNA testing or parentage. Smoky black has been mistaken for faded black, dark bay or brown, grullo or even liver chestnut.

<span class="mw-page-title-main">Labrador Retriever coat colour genetics</span> Genetics behind Labrador Retriever coat colour

The genetic basis of coat colour in the Labrador Retriever has been found to depend on several distinct genes. The interplay among these genes is used as an example of epistasis.

<span class="mw-page-title-main">Amelanism</span> Pigmentation abnormality

Amelanism is a pigmentation abnormality characterized by the lack of pigments called melanins, commonly associated with a genetic loss of tyrosinase function. Amelanism can affect fish, amphibians, reptiles, birds, and mammals including humans. The appearance of an amelanistic animal depends on the remaining non-melanin pigments. The opposite of amelanism is melanism, a higher percentage of melanin.

<span class="mw-page-title-main">Seal brown (horse)</span> Hair coat color of horses

Seal brown is a hair coat color of horses characterized by a near-black body color; with black points, the mane, tail and legs; but also reddish or tan areas around the eyes, muzzle, behind the elbow and in front of the stifle. The term is not to be confused with "brown", which is used by some breed registries to refer to either a seal brown horse or to a dark bay without the additional characteristics of seal brown.

<span class="mw-page-title-main">Melanistic mask</span> Dog coat pattern

A melanistic mask is a dog coat pattern that gives the appearance of a mask on the dog's face. The hairs on the muzzle, and sometimes entire face or ears, are colored by eumelanin instead of pheomelanin pigment. Eumelanin is typically black, but may instead be brown, dark gray, or light gray-brown. Pheomelanin ranges in color from pale cream to mahogany. The trait is caused by M264V (EM), a completely dominant allele (form) of the melanocortin 1 receptor gene.

<span class="mw-page-title-main">Dog coat genetics</span> Genetics behind dog coat

Dogs have a wide range of coat colors, patterns, textures and lengths. Dog coat color is governed by how genes are passed from dogs to their puppies and how those genes are expressed in each dog. Dogs have about 19,000 genes in their genome but only a handful affect the physical variations in their coats. Most genes come in pairs, one being from the dog's mother and one being from its father. Genes of interest have more than one expression of an allele. Usually only one, or a small number of alleles exist for each gene. In any one gene locus a dog will either be homozygous where the gene is made of two identical alleles or heterozygous where the gene is made of two different alleles.

References

  1. Maerz and Paul A Dictionary of Color New York: 1930 McGraw-Hill Page 198; Color Sample of Liver: Page 37 Plate 7 Color Sample H9.
  2. Animal Genetics. "B-Locus (Brown, Liver, Chocolate)". www.horsetesting.com. Retrieved 2014-05-05.Use of copyrighted material rationale: no material was used, just referenced.
  3. Henner, J; PA Poncet; L Aebi; C Hagger; G Stranzinger; S Rieder (August 2002). "Horse breeding: genetic tests for the coat colors chestnut, bay and black. Results from a preliminary study in the Swiss Freiberger horse breed". Schweizer Archiv für Tierheilkunde. 144 (8): 405–412. doi:10.1024/0036-7281.144.8.405. PMID   12224446. The statistical analysis of 1369 offspring from five stallions indicate, that darker shades of basic color phenotypes (dark chestnut, dark bay) follow a recessive mode of inheritance in the Franches-Montagnes horse breed.
  4. Strassburg, Christian P. (2010). "Hyperbilirubinemia syndromes (Gilbert-Meulengracht, Crigler-Najjar, Dubin-Johnson, and Rotor syndrome)". Best Practice & Research: Clinical Gastroenterology. 24 (5): 555–571. doi:10.1016/j.bpg.2010.07.007. PMID   20955959.
  5. Unknown, Various (Spring 2012). "Hepatic Pathology Index". Mercer School of Medicine Internet Pathology Laboratory for Medical Education. Retrieved 2013-01-21.Fair Use Rationale: this is an educational journal.