Pinto horse

Last updated

A pinto horse, with patches of white and of another color P1320185 - Flickr - gailhampshire.jpg
A pinto horse, with patches of white and of another color

A pinto horse has a coat color that consists of large patches of white and any other color. Pinto coloration is also called paint, [1] particolored, [2] or in nations that use British English, simply coloured. Pinto horses have been around since shortly after the domestication of the horse. [3]

Contents

Pinto colors can come in a number of genetically distinct patterns, which have different visual characteristics and tend to make white or leave colored different areas of the horse. These include tobiano, sabino, splashed white, frame, and manchado. A pinto horse may also have a combination of these patterns, such as tovero. [2]

Pinto patterns can be found in various breeds of horses, notably including the American Paint Horse. Color breed registries such as the Pinto Horse Association of America record pedigree and horse show results for pinto horses, regardless of ancestry. Both the terms "Pinto" and "Paint" may sometimes refer to breeds or registries rather than coat color. [2]

Pinto patterns are visually and genetically distinct from the leopard complex spotting patterns characteristic of horse breeds such as the Appaloosa. Breeders who select for color are often careful not to cross the two patterns, and registries that include spotting color preferences often refuse registration to horses that exhibit characteristics of the "wrong" pattern.

Description

A pinto horse has a coat with patches of white fur and patches of another color. The white on a pinto horse is generally asymmetric, unlike for example white added by the leopard complex. [2]

The non-white area has the same colors in the same arrangements as one would see on a solid horse. Overall, the effect is as if a horse with a solid coat had white painted in patches over top. The white areas of a pinto horse generally have pink skin underneath.

A horse with small amounts of white only on the face and/or legs is not called "pinto" but instead said to have white markings. There is no clear dividing line for how much white counts as pinto and how much counts as only white markings, and various breed registries have slightly different rules on how much white must be present and where it must be placed to count as pinto. [4] [5] [6]

The word pinto is Spanish for "painted", "dappled", or "spotted". [7]

History

A horse with tobiano-like spotting depicted on a Corinthian black-figure column krater from 570-560 BC Amphiaraos Krater chariot race cropped.png
A horse with tobiano-like spotting depicted on a Corinthian black-figure column krater from 570-560 BC

The earliest known pinto horses appeared shortly after horses were domesticated. Analysis of ancient horse DNA found a tobiano horse that lived about 5600 years ago, [8] and a sabino-1 horse that lived about 5000 years ago. [3] Tobiano was at first favored by humans and became especially common during the Iron Age (900 BC to 400 AD), before becoming less frequent again in the Middle Ages. [8]

Images from pottery and other art of ancient antiquity show horses with flashy, spotted patterns, indicating that they may have been desirable traits and selectively bred for. Images of spotted horses appear in the art of Ancient Egypt, and archaeologists have found evidence of horses with spotted coat patterns on the Russian steppes before the rise of the Roman Empire. Later, spotted horses were among those brought to the Americas by the conquistadores .

By the 17th century in Europe, spotted horses were quite fashionable, though when the fad ended, large numbers of newly unsellable horses were shipped to the Americas, some of which were sold, while others were simply turned loose to run wild. [9] The color became popular, particularly among Native Americans, and was specifically bred for in the United States, which now has the greatest number of pinto horses in the world.

Pinto coats by base color

Piebald horse - geograph.org.uk - 1317208.jpg
Piebald
Icelandic Horse MichaD.jpg
Skewbald
Tobiano.jpg
Tricolored

A few words describe pinto horses by giving more detail about the color of the non-white areas, mainly used in British English. This can also be done by including the base color in the coat name, such as "bay pinto" or "pinto palomino".

Pinto coats by pattern

While pinto horses in general have patches of white and patches of color, there are a number of different, separately inherited patterns which tend to arrange the white and colored areas differently.

PatternDescription
Horse at Ocracoke Pony Pen by Bonnie Gruenberg.jpg Tobiano The most recognizable type of pinto, tobiano is a spotting pattern characterized by rounded markings with white legs and white across the back between the withers and the dock of the tail, usually arranged in a roughly vertical pattern. The head is usually dark, and may have facial markings similar to those seen on non-pinto horses such as a star, snip, strip, or blaze. Tobiano is a dominant trait caused by a single gene, so all tobiano horses have at least one tobiano parent. [10]
Overo paint horse by Bonnie Gruenberg.jpg Frame or frame overo Frame is characterized by horizontally oriented white patches with jagged, crisp edges. White patches typically include the head, face, and lateral aspects of the neck and body, and the eyes can be blue. Frame overos may have very modest markings that are not obviously pinto. This quality allows the pattern to seemingly "hide" for generations, and is thought to be responsible for some cases of cropouts.

The same gene that causes the frame pattern can also cause lethal white syndrome. Although frame overo horses are themselves healthy, if two horses with the frame gene are bred together, there is a 25% chance that the foal will have lethal white syndrome. Affected foals are fully white and die shortly after birth. [11]

Splashwhitemare2.jpg Splashed white A less-common pinto pattern, splashed white coats have horizontally oriented white markings with crisp, smooth edges, and make the horse appear to have been dipped, head lowered, into white paint. The face has significant white markings, and the eyes are usually blue. Most splashed white pintos have normal hearing, but the trait is linked to congenital deafness. Splashed white patterns can be caused by multiple variants of two different genes. [12]
Sato - Palomino sabino Purebred Thoroughbred Stallion (5966320009).jpg Sabino and dominant white A group of visually similar patterns which may range from white markings on the face and legs all the way up to a solid white horse. In between those extremes, sabino horses possess a spotting pattern characterized by high white on legs, belly spots, and white markings on the face extending past the eyes. The edges of markings may be "lacy" or patches of roaning patterns standing alone or on the edges of white markings can occur. Sabino may sometimes look similar to roan or rabicano. Of the 30+ forms of sabino and dominant white where the genetic cause is known, all are caused by different alternate variants of the same gene. It's possible that some forms of sabino where the cause is not yet known may turn out to be caused by a different gene or genes. [2]
ManchadoA rare pattern characterized by large, crisp white areas with smooth round spots of color inside them. Usually there will be more white on the dorsal side the neck, and more color on the belly and the ventral side of the neck. It has been seen in the Thoroughbred, Criollo, Polo Pony, Arabian, and Hackney breeds, and almost always in Argentina. The cause of manchado is not known for certain, but Sponenberg & Bellone propose it may be caused by a rare recessive gene. [2]

Some additional terms describe the pattern without specifying the exact pinto pattern. These include:

A "medicine hat" marking, dark ears on a white head APHA Mare2.jpg
A "medicine hat" marking, dark ears on a white head

In biology, pinto is considered a type of piebaldism, which is itself a type of leucism.

Lethal white syndrome

As noted in the description of patterns, the frame gene is associated with a condition called lethal white syndrome. This is a genetic disorder causing foals to die shortly after birth. Affected foals are fully white and have a non-functional colon. [11]

The gene that causes lethal white syndrome is the same gene that causes the frame overo pinto pattern. A single copy results in a frame overo horse, while two copies being present causes lethal white syndrome. Some horses may have the gene without visually appearing to be frame patterned, but a DNA test exists to determine whether a horse is a carrier. Lethal white syndrome can be avoided by not breeding two carriers together. [11]

Organizations

Pinto (left), leopard-spotted Appaloosa (right) Gedinne 050717 (1).JPG
Pinto (left), leopard-spotted Appaloosa (right)

A number of color breed registries encourage the breeding of pinto-colored horses, with varying registration requirements. The less restrictive organizations allow registration of a horse of any breed or combination of breeds with as little as three square inches of white above the knees or hocks, not including facial markings. Some pinto registries do not accept animals with draft horse or mule breeding, though others do. None accepts horses with the genetically distinct Appaloosa pattern, produced by genes in the leopard complex, and the Appaloosa registry in turn does not accept animals with pinto patterns.

When used to refer to breeds, Pinto is a color breed that can be of any type or ancestry, while a Paint is a breed with a specific type and bloodlines.

Pinto Horse Association of America

The Pinto Horse Association of America (PtHA) is a color breed, which accepts horses based on coat color regardless of ancestry. They maintain three registries: the Color Registry, which accepts pinto colored horses; the Solid Registry, which accepts any horse not accepted by the Colored Registry; and the Long Ear Registry, which accepts donkeys and mules of any color. To count as Pinto, a horse must have at least four square inches of white fur and pink skin in areas other than the face and lower legs. The Color Registry does not accept horses with appaloosa characteristics, but the Solid Registry does. The Color Registry also requires that both parents of stallions be registered with the PtHA or an approved outcross breed, though this is not required for ponies, miniatures, or horses of Vanner or Drum type. [6] [15] [16]

The PtHA has over 157,000 registered Pintos. [17]

American Paint Horse Association

Horses with pinto coloring and verifiable pedigrees tracing to Quarter Horses or Thoroughbreds have been named the American Paint Horse, and are recorded in a separate registry, the American Paint Horse Association. While a Pinto may be of any breed or combination of breeds (possibly with restrictions depending on the registry), a horse that is registered as an American Paint Horse must have at least one parent recorded with the APHA, and both parents must be only of registered American Quarter Horse, American Paint Horse, or Thoroughbred bloodlines. Therefore, most Paint horses may also be registered as Pintos, but not all Pintos are qualified to be registered as Paints.

The American Paint Horse is a very popular breed in the United States, with around 10,000 horses registered annually, roughly two thirds of which are in the US. [18]

Pinto in other breeds

Many breed registries do not, or at some point in the past did not, accept cropout horses with spots or "excess" white for registration, believing that such animals were likely to be crossbreds, or due to a fear of producing lethal white foals. This exclusion of offspring from pedigreed parents led to the formation not only of the American Paint Horse Association, but also other pinto registries. Among the breeds that excluded such horses were the Arabian Horse and American Quarter Horse registries. Modern DNA testing, though, has revealed that some breeds do possess genes for spotting patterns, such as a non-SB-1 sabino pattern in Arabians, and sabino, overo, and tobiano in Quarter Horses. Therefore, these registries have modified their rules, allowing horses with extra white, if parentage is verified through DNA testing, to be registered. The Jockey Club's Thoroughbred registry, however, still does not officially recognize pinto as a registerable color, though it does allow white body spots to be recorded under the category of markings. [19] The Welsh Pony and Cob Society of the UK also does not accept "piebald" or "skewbald" horses for registration. [20]

See also

Related Research Articles

<span class="mw-page-title-main">Appaloosa</span> American horse breed noted for spotted color pattern

The Appaloosa is an American horse breed best known for its colorful spotted coat pattern. There is a wide range of body types within the breed, stemming from the influence of multiple breeds of horses throughout its history. Each horse's color pattern is genetically the result of various spotting patterns overlaid on top of one of several recognized base coat colors. The color pattern of the Appaloosa is of interest to those who study equine coat color genetics, as it and several other physical characteristics are linked to the leopard complex mutation (LP). Appaloosas are prone to develop equine recurrent uveitis and congenital stationary night blindness; the latter has been linked to the leopard complex.

<span class="mw-page-title-main">Roan (color)</span>

Roan is a coat color found in many animals, including horses, cattle, antelope, cat and dogs. It is defined generally as an even mixture of white and pigmented hairs that do not "gray out" or fade as the animal ages. There are a variety of genetic conditions which produce the colors described as "roan" in various species.

A color breed refers to groupings of horses whose registration is based primarily on their coat color, regardless of the horse's actual breed or breed type.

<span class="mw-page-title-main">American Paint Horse</span> American breed of horse

The American Paint Horse is a breed of horse that combines both the conformational characteristics of a western stock horse with a pinto spotting pattern of white and dark coat colors. Developed from a base of spotted horses with Quarter Horse and Thoroughbred bloodlines, the American Paint Horse Association (APHA) breed registry is now one of the largest in North America. The registry allows some non-spotted animals to be registered as "Solid Paint Bred" and considers the American Paint Horse to be a horse breed with distinct characteristics, not merely a color breed.

<span class="mw-page-title-main">Lethal white syndrome</span> Genetic disorder in horses

Lethal white syndrome (LWS), also called overo lethal white syndrome (OLWS), lethal white overo (LWO), and overo lethal white foal syndrome (OLWFS), is an autosomal genetic disorder most prevalent in the American Paint Horse. Affected foals are born after the full 11-month gestation and externally appear normal, though they have all-white or nearly all-white coats and blue eyes. However, internally, these foals have a nonfunctioning colon. Within a few hours, signs of colic appear; affected foals die within a few days. Because the death is often painful, such foals are often humanely euthanized once identified. The disease is particularly devastating because foals are born seemingly healthy after being carried to full term.

<span class="mw-page-title-main">Piebald</span> Animal with white markings on a darker coat

A piebald or pied animal is one that has a pattern of unpigmented spots (white) on a pigmented background of hair, feathers or scales. Thus a piebald black and white dog is a black dog with white spots. The animal's skin under the white background is not pigmented.

<span class="mw-page-title-main">Skewbald</span>

Skewbald is a colour pattern of horses. A skewbald horse has a coat made up of white patches on a non-black base coat, such as chestnut, bay, or any colour besides black coat. Skewbald horses which are bay and white are sometimes called tricoloured. These horses usually have pink skin under white markings and dark skin under non-white areas. Other than colour, it is similar in appearance to the piebald pattern. Some animals also exhibit colouration of the irises of the eye that match the surrounding skin. The underlying genetic cause is related to a condition known as leucism. The term is also used to describe spotting patterns in various other animals, such as goats.

<span class="mw-page-title-main">Tricoloured horse</span>

Tri-coloured refers to a horse with three different coat colours in a pinto spotting pattern of large white and dark patches, usually bay and white. This colouring is also commonly called skewbald. In modern usage in British English, skewbald and piebald horses are collectively referred to as coloured, while in North American English, the term pinto is used to describe the colour pattern.

<span class="mw-page-title-main">Tobiano</span> Spotted color pattern in horses

Tobiano is a spotted color pattern commonly seen in pinto horses, produced by a dominant gene. The tobiano gene produces white-haired, pink-skinned patches on a base coat color. The coloration is almost always present from birth and does not change throughout the horse's lifetime, unless the horse also carries the gray gene. It is a dominant gene, so any tobiano horse must have at least one parent who carries the tobiano gene.

<span class="mw-page-title-main">Overo</span> Group of colouration patterns of horses

Overo refers to several genetically unrelated pinto coloration patterns of white-over-dark body markings in horses, and is a term used by the American Paint Horse Association to classify a set of pinto patterns that are not tobiano. Overo is a Spanish word, originally meaning "like an egg". The most common usage refers to frame overo, but splashed white and sabino are also considered "overo". A horse with both tobiano and overo patterns is called tovero.

<span class="mw-page-title-main">Equine coat color</span> Horse coat colors and markings

Horses exhibit a diverse array of coat colors and distinctive markings. A specialized vocabulary has evolved to describe them.

<span class="mw-page-title-main">Sabino horse</span> Color pattern in horses

Sabino describes a distinct pattern of white spotting in horses. In general, Sabino patterning is visually recognized by roaning or irregular edges of white markings, belly spots, white extending past the eyes or onto the chin, white above the knees or hocks, and "splash" or "lacy" marks anywhere on the body. Some sabinos have patches of roan patterning on part of the body, especially the barrel and flanks. Some sabinos may have a dark leg or two, but many have four white legs. Sabino patterns may range from slightly bold face or leg white markings—as little as white on the chin or lower lip—to horses that are fully white.

<span class="mw-page-title-main">Horse markings</span> Colored areas, usually white, on a horse that differ from the body color

Markings on horses are usually distinctive white areas on an otherwise dark base coat color. Most horses have some markings, and they help to identify the horse as a unique individual. Markings are present at birth and do not change over the course of the horse's life. Most markings have pink skin underneath most of the white hairs, though a few faint markings may occasionally have white hair with no underlying pink skin. Markings may appear to change slightly when a horse grows or sheds its winter coat, however this difference is simply a factor of hair coat length; the underlying pattern does not change.

The American Paint Horse Association (APHA) is a breed registry for the American Paint Horse. It is currently headquartered in Fort Worth, Texas. It was founded in 1965 with the merging of two different color breed registries that had been formed to register pinto-colored horses of Quarter Horse bloodlines. One of these organizations was the American Paint Quarter Horse Association and the other was the American Paint Stock Horse Association.

A cropout, crop-out or crop out is a horse with body spots, including pinto or leopard complex spotting, or "high white" horse markings, with a sire and dam who both appeared to have been solid-colored. There are several variations in the definition, depending on the breed registry involved. There are multiple genetic reasons that may cause a horse to be a cropout. Sometimes solid-colored horses throw cropouts because some spotting patterns are recessive genes that are not necessarily expressed unless the relevant allele is inherited from both parents. Other times a gene may be a dominant or incomplete dominant but so minimally expressed that the horse looks solid but can pass flashy color on to its offspring.

<span class="mw-page-title-main">Splashed white</span>

Splashed white or splash is a horse coat color pattern in the "overo" group of spotting patterns that produces pink-skinned, white markings. Many splashed whites have very modest markings, while others have the distinctive "dipped in white paint" pattern. Blue eyes are a hallmark of the pattern, and splash may account for otherwise "solid" blue-eyed horses. Splashed white occurs in a variety of geographically divergent breeds, from Morgans in North America to Kathiawari horses in India. The splashed white pattern is also associated with congenital deafness, though most splashed whites have normal hearing. Splashed white can be caused by multiple variants across two different genes, for which genetic testing is available.

<span class="mw-page-title-main">Dominant white</span> Horse coat color and its genetics

Dominant white (W) is a group of genetically related coat color alleles on the KIT gene of the horse, best known for producing an all-white coat, but also able to produce various forms of white spotting, as well as bold white markings. Prior to the discovery of the W allelic series, many of these patterns were described by the term sabino, which is still used by some breed registries.

The Pinto Horse Association of America (PtHA) registers horses, utility horses, ponies and miniature horses of various pedigrees with certain kinds of pinto coat colors. The word pinto is Spanish for "paint." In general terms, pinto can apply to any horse marked with unpigmented pink-skinned, white-haired areas on its coat. The Pinto Horse Association of America provides the owners and riders of pintos with a show circuit and a breed organization. The primary requirement for PtHA registration is coat color; the pinto is not a true breed, but a color breed.

<span class="mw-page-title-main">Roan (horse)</span> Horse coat color pattern characterized by an even mixture of colored and white hairs on the body

Roan is a horse coat color pattern characterized by an even mixture of colored and white hairs on the body, while the head and "points"—lower legs, mane, and tail—are mostly solid-colored. Horses with roan coats have white hairs evenly intermingled throughout any other color. The head, legs, mane, and tail have fewer scattered white hairs or none at all. The roan pattern is dominantly inherited, and is found in many horse breeds. While the specific mutation responsible for roan has not been exactly identified, a DNA test can determine zygosity for roan in several breeds. True roan is always present at birth, though it may be hard to see until after the foal coat sheds out. The coat may lighten or darken from winter to summer, but unlike the gray coat color, which also begins with intermixed white and colored hairs, roans do not become progressively lighter in color as they age. The silvering effect of mixed white and colored hairs can create coats that look bluish or pinkish.

<span class="mw-page-title-main">Pintabian</span> Breed of horse

The Pintabian horse is an American part-Arabian horse breed. It has over 99% Arabian blood, but also exhibits the tobiano color pattern, which is not found in thoroughbred Arabians. The registry began using the term "Pintabian" in 1992 and trademarked the word in 1995. which is the official registering authority for Pintabian horses worldwide.

References

  1. "A Guide to Equine Color Genetics and Coat Color". Practical Horseman. Retrieved January 1, 2024.
  2. 1 2 3 4 5 6 7 Sponenberg, D. Phillip; Bellone, Rebecca (2017). Equine Color Genetics (4 ed.). Wiley Blackwell. pp. 171, 191, 195–196.
  3. 1 2 Ludwig, Arne; Pruvost, Melanie; Reissmann, Monika; Benecke, Norbert; Brockmann, Gudrun A.; Castaños, Pedro; Cieslak, Michael; Lippold, Sebastian; Llorente, Laura; Malaspinas, Anna-Sapfo; Stalkin, Montgomery; Hofreiter, Michael (2009). "Coat Color Variation at the Beginning of Horse Domestication". Science. 324 (5926): 485. Bibcode:2009Sci...324..485L. doi:10.1126/science.1172750. PMC   5102060 . PMID   19390039.
  4. "2024 AQHA Rulebook". American Quarter Horse Association. Retrieved January 1, 2024.
  5. "2023 Official APHA Rule Book" (PDF). American Paint Horse Association. Retrieved January 1, 2024.
  6. 1 2 "Color Breed Registry". Pinto Horse Association of America. Retrieved January 1, 2024.
  7. "pinto". WordReference.com Spanish-English Dictionary. Retrieved October 27, 2012.
  8. 1 2 Wutke, Saskia; Benecke, Norbert; Sandoval-Castellanos, Edson; Döhle, Hans-Jürgen; Friederich, Susanne; Gonzalez, Javier; Hallsson, Jón Hallsteinn; Hofreiter, Michael; Lõugas, Lembi; Magnell, Ola; Morales-Muniz, Arturo; Orlando, Ludovic; Pálsdóttir, Albína Hulda; Reissmann, Monika; Ruttkay, Matej; Trinks, Alexandra; Ludwig, Arne (December 23, 2016). "Spotted phenotypes in horses lost attractiveness in the Middle Ages". Scientific Reports. 6 (1): 38548. Bibcode:2016NatSR...638548W. doi:10.1038/srep38548. PMC   5141471 . PMID   27924839.
  9. Bennett, Deb. Conquerors: The Roots of New World Horsemanship. Amigo Publications Inc; 1st edition 1998. ISBN   0-9658533-0-6.
  10. "Tobiano - Horse Coat Color". www.vgl.ucdavis.edu. Archived from the original on May 15, 2010. Retrieved November 18, 2017.
  11. 1 2 3 Metallinos, DL; Bowling AT; Rine J (June 1998). "A missense mutation in the endothelin-B receptor gene is associated with Lethal White Foal Syndrome: an equine version of Hirschsprung Disease". Mammalian Genome. 9 (6). New York: Springer New York: 426–31. doi:10.1007/s003359900790. PMID   9585428. S2CID   19536624.
  12. "Splashed White Mutations". UC Davis VGL. Retrieved January 1, 2024.
  13. Ashley Stollberg. "Etching a Pinto Pattern". Breyer. Retrieved December 31, 2023.
  14. Leah Posey-Patton. "Seeing Spots: Pinto Patterns in Equines". Breyer. Retrieved December 31, 2023.
  15. "Solid Registry". Pinto Horse Association of America. Retrieved January 1, 2024.
  16. "Long Ear Registry". Pinto Horse Association of America. Retrieved January 1, 2024.
  17. "Home". Pinto Horse Association of America. Retrieved January 1, 2024.
  18. "2022 Annual Report" (PDF). American Paint Horse Association. Retrieved January 1, 2024.
  19. "Jockey Club Interactive Registration". www.registry.jockeyclub.com. Retrieved November 18, 2017.
  20. "Registration Regulations". wpcs.uk.com. Retrieved November 18, 2017.