Heterogyrus

Heterogyrus milloti
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Family:	Gyrinidae
Subfamily:	Heterogyrinae
Genus:	Heterogyrus ; Legros, 1953
Species:	H. milloti
Binomial name
	Heterogyrus milloti; Legros, 1953

Last updated January 03, 2026

Heterogyrus milloti is a species of beetle in the family Gyrinidae, the only species in the genus Heterogyrus.^[1] It is endemic to Madagascar.

Within Gyrinidae, the genus Spanglerogyrus is sister to all other taxa; Heterogyrus is, in turn, sister to all gyrinids except Spanglerogyrus, with an estimated divergence 200 million years ago.^[2]^[3]^[4] It is suggested to be closely related to the fossil genera Mesogyrus, Cretotortor and Baissogyrus which constitute the subfamily Heterogyrinae.^[2]

References

↑ Gyrinidae Species List Archived 2012-10-23 at the Wayback Machine at Joel Hallan's Biology Catalog. Texas A&M University. Retrieved on 9 May 2012.
1 2 Gustafson, Grey T.; Prokin, Alexander A.; Bukontaite, Rasa; Bergsten, Johannes; Miller, Kelly B. (2017-08-22). "Tip-dated phylogeny of whirligig beetles reveals ancient lineage surviving on Madagascar". Scientific Reports. 7 (1): 8619. Bibcode:2017NatSR...7.8619G. doi: 10.1038/s41598-017-08403-1 . ISSN 2045-2322. PMC 5567340 . PMID 28831048.
↑ Miller, Kelly B.; Bergsten, Johannes (October 2012). "Phylogeny and classification of whirligig beetles (Coleoptera: Gyrinidae): relaxed‐clock model outperforms parsimony and time‐free Bayesian analyses". Systematic Entomology. 37 (4): 706–746. doi:10.1111/j.1365-3113.2012.00640.x. ISSN 0307-6970.
↑ Beutel, Rolf G.; Yan, Evgeny; Yavorskaya, Margarita; Büsse, Sebastian; Gorb, Stanislav N.; Wipfler, Benjamin (April 2019). "On the thoracic anatomy of the Madagascan Heterogyrus milloti and the phylogeny of Gyrinidae (Coleoptera)". Systematic Entomology. 44 (2): 336–360. doi:10.1111/syen.12325. ISSN 0307-6970.

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[1] Gyrinidae Species List Archived 2012-10-23 at the Wayback Machine at Joel Hallan's Biology Catalog. Texas A&M University. Retrieved on 9 May 2012.

[:0-2] 1 2 Gustafson, Grey T.; Prokin, Alexander A.; Bukontaite, Rasa; Bergsten, Johannes; Miller, Kelly B. (2017-08-22). "Tip-dated phylogeny of whirligig beetles reveals ancient lineage surviving on Madagascar". Scientific Reports. 7 (1): 8619. Bibcode:2017NatSR...7.8619G. doi: 10.1038/s41598-017-08403-1 . ISSN 2045-2322. PMC 5567340 . PMID 28831048.

[3] Miller, Kelly B.; Bergsten, Johannes (October 2012). "Phylogeny and classification of whirligig beetles (Coleoptera: Gyrinidae): relaxed‐clock model outperforms parsimony and time‐free Bayesian analyses". Systematic Entomology. 37 (4): 706–746. doi:10.1111/j.1365-3113.2012.00640.x. ISSN 0307-6970.

[4] Beutel, Rolf G.; Yan, Evgeny; Yavorskaya, Margarita; Büsse, Sebastian; Gorb, Stanislav N.; Wipfler, Benjamin (April 2019). "On the thoracic anatomy of the Madagascan Heterogyrus milloti and the phylogeny of Gyrinidae (Coleoptera)". Systematic Entomology. 44 (2): 336–360. doi:10.1111/syen.12325. ISSN 0307-6970.

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