Hiya (plant)

Hiya
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Family:	Dennstaedtiaceae
Genus:	Hiya ; H.Shang
Type species
	Hiya brooksiae; (Alderw.) H.Shang

Last updated October 01, 2025

Hiya is a genus of ferns belonging to the family Dennstaedtiaceae. Described in 2018, it resembles ferns of the genus Hypolepis but differs from it by multiple characteristics: scrambling, indeterminate and intermittent growth of fronds; stipule-like pinnules at the base of pinnae, and a rachis-costa architecture where the adaxial sulcus of the rachis is continuous with that of the costae and costules.^[1]^[2]

The name of the genus references the name given to imperial guards from the Qing dynasty of China, hiya, due to the prickly or very rough armed stipes present in the ferns.^[1]

Classification

As of January 2023^[update], the following species are accepted in the genus as currently circumscribed by the Checklist of Ferns and Lycophytes of the World:^[3]

Hiya brooksiae (Alderw.) H.Shang (= Hypolepis brooksiaeAlderw. = Hypolepis celebicaC. Chr.)
Hiya distans (Hook.) Brownsey & Perrie (= Hypolepis distansHook.)
Hiya nigrescens (Hook.) H.Shang (= Hypolepis nigrescensHook. = Hypolepis hispanolicaMaxon = Dennstaedtia rubicaulisChrist)
Hiya scabristipes (Brownsey) H.Shang (= Hypolepis scabristipesBrownsey)

References

1 2 3 Shang H, Sundue M, Wei R, Wei X, Luo J, Liu L, et al. (2018). "Hiya: A new genus segregated from Hypolepis in the fern family Dennstaedtiaceae, based on phylogenetic evidence and character evolution". Mol Phylogenet Evol. 127: 449–458. doi:10.1016/j.ympev.2018.04.038. PMID 29723646. S2CID 19089119.
↑ Schwartsburd PB, Perrie LR, Brownsey P, Shepherd LD, Shang H, Barrington DS, et al. (2020). "New insights into the evolution of the fern family Dennstaedtiaceae from an expanded molecular phylogeny and morphological analysis". Mol Phylogenet Evol. 150 106881. doi:10.1016/j.ympev.2020.106881. ISSN 1055-7903. PMID 32512193. S2CID 219550667.
↑ Hassler, Michael (19 January 2023). "Hiya". Checklist of Ferns and Lycophytes of the World. 14.1. Retrieved January 21, 2023.

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[Hiya_fern-1] 1 2 3 Shang H, Sundue M, Wei R, Wei X, Luo J, Liu L, et al. (2018). "Hiya: A new genus segregated from Hypolepis in the fern family Dennstaedtiaceae, based on phylogenetic evidence and character evolution". Mol Phylogenet Evol. 127: 449–458. doi:10.1016/j.ympev.2018.04.038. PMID 29723646. S2CID 19089119.

[Insights_into_Dennstaedtiaceae-2] Schwartsburd PB, Perrie LR, Brownsey P, Shepherd LD, Shang H, Barrington DS, et al. (2020). "New insights into the evolution of the fern family Dennstaedtiaceae from an expanded molecular phylogeny and morphological analysis". Mol Phylogenet Evol. 150 106881. doi:10.1016/j.ympev.2020.106881. ISSN 1055-7903. PMID 32512193. S2CID 219550667.

[CFLW-3] Hassler, Michael (19 January 2023). "Hiya". Checklist of Ferns and Lycophytes of the World. 14.1. Retrieved January 21, 2023.

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