Hollardosteus Temporal range: Eifelian | |
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reconstruction as a dunkleosteid | |
Scientific classification | |
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Genus: | Hollardosteus |
Species: | H. marocanus |
Binomial name | |
Hollardosteus marocanus Lehman, 1976 | |
Hollardosteus marocanus is a brachythoracid arthrodire placoderm from the Middle Devonian of Morocco. Its scrappy remains were found in Eifelian-aged strata from the Tafilat-Maider Valley. [1]
The inferognathal plate has a well-developed anterior tooth (prong), very similar in form to the inferognathals of the dunkleosteids. However, the dermal surface of the armor is decorated with a pattern of large, crowded tubercules totally unlike the smooth dermal surfaces of dunkleosteid armors. This mix of peculiar features leaves H. marocanus placed as "Arthrodira incertae sedis," [1] at least until more complete specimens are found.
Chondrichthyes is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fishes, which have skeletons primarily composed of bone tissue. Chondrichthyes are jawed vertebrates with paired fins, paired nares, scales, and a heart with its chambers in series. Extant chondrichthyes range in size from the 10 cm finless sleeper ray to the over 10 m whale shark.
In most biological nomenclature, a scale is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran species, scales are plates on the surface of the insect wing, and provide coloration. Scales are quite common and have evolved multiple times through convergent evolution, with varying structure and function.
The dermis or corium is a layer of skin between the epidermis and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided into two layers, the superficial area adjacent to the epidermis called the papillary region and a deep thicker area known as the reticular dermis. The dermis is tightly connected to the epidermis through a basement membrane. Structural components of the dermis are collagen, elastic fibers, and extrafibrillar matrix. It also contains mechanoreceptors that provide the sense of touch and thermoreceptors that provide the sense of heat. In addition, hair follicles, sweat glands, sebaceous glands, apocrine glands, lymphatic vessels, nerves and blood vessels are present in the dermis. Those blood vessels provide nourishment and waste removal for both dermal and epidermal cells.
Crocodile armor consists of the protective dermal and epidermal components of the integumentary system in animals of the order Crocodilia.
The class Osteostraci is an extinct taxon of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian.
Platyhystrix is an extinct temnospondyl amphibian with a distinctive sail along its back, similar to the unrelated synapsids, Dimetrodon and Edaphosaurus. It lived during the boundary between the latest Carboniferous and earliest Permian periods throughout what is now known as the Four Corners, Texas, and Kansas about 300 million years ago.
Cyamodus is a genus of placodonts known from several species described from Middle-Late Triassic of Europe and China. The genus was described by Christian Erich Hermann von Meyer in 1863, based on specimens found in Germany. Like some other placodonts, Cyamodus has an armoured carapace composed of irregular hexagonal plates, with the mouth containing a small number of large, rounded teeth that were likely involved in crushing hard shelled organisms (durophagy).
Phyllolepis is the type genus of Phyllolepida, an extinct taxon of arthrodire placoderm fish from the middle to late Devonian. The species of Phyllolepis, themselves, are restricted to the Famennian-aged freshwater strata of the Late Devonian, around 360 million years ago. Fossils of this genus have been found primarily in Europe and North America. The end of the Devonian saw them disappear in a mass extinction.
Laidleria is an extinct genus of temnospondyl that likely lived between the Early to Middle Triassic, though its exact stratigraphic range is less certain. Laidleria has been found in the Karoo Basin in South Africa, in Cynognathus Zone A or B. The genus is represented by only one species, L. gracilis, though the family Laidleriidae does include other genera, such as Uruyiella, sister taxon to Laidleria, which was discovered and classified in 2007.
Holonema is an extinct genus of relatively large, barrel-shaped arthrodire placoderms that were found in oceans throughout the world from the Mid to Late Devonian, when the last species perished in the Frasnian-Fammian extinction event. Most species of the genus are known from fragments of their armor, but the Gogo Reef species, H. westolli, is known from whole, articulated specimens.
Murrindalaspis is an extinct genus of acanthothoracid placoderm found in the McLarty Member of the Murrindal Limestone, of the Early Devonian-aged Buchan Group in eastern Victoria, Australia. Murrindalaspis differs from other acanthothoracids by having a dorsal plate with a large, blade-like flattened, recurved crest emanating from the medial line, and no ventral keel. So far, the genus is known only from dorsal plates and ossified eyeballs. The genus differs from the closely related Weejasperaspis in that the dorsal crest of the latter is shorter, and triangular-shaped.
Dissorophus (DI-soh-ROH-fus) is an extinct genus of temnospondyl amphibian that lived during the Early Permian Period about 273 million years ago. Its fossils have been found in Texas and in Oklahoma in North America. Its heavy armor and robust build indicate Dissorophus was active on land, similar to other members of the clade Dissorophidae that are known from the Late Carboniferous to the Early Permian periods. Dissorphus is distinguished by its small body size, disproportionately large head and short trunk.
Psephosauriscus is an extinct genus of placodont reptile from the Middle Triassic of Israel and Egypt. It is known from bony armor plates that have been found from Makhtesh Ramon in Israel's Negev desert and Araif en Naqua on Egypt's Sinai Peninsula. The genus was erected in 2002 as a replacement name for several species of the genus Psephosaurus, which was named in 1957. It includes the species P. mosis, P. ramonensis, P. sinaiticus, and a possible fourth species, P. rhombifer. All species, with the exception of P. ramonensis, were once assigned to the genus Psephosaurus. Remains of P. mosis and P. ramonensis were found in Makhtesh Ramon, while P. sinaiticus and P. rhombifer were found in Araif en Naqua.
Holonematidae is an extinct family of relatively large arthrodire placoderms from the Early to Late Devonian. Almost all fossil specimens are of armor fragments, though, all have distinctive ornamentation, often of unique arrangements and patterns of tubercles, that are diagnostic of the family. The trunkshield is very elongated, giving the armor an overall "barrel" like appearance.
Buchanosteidae is a family of arthrodire placoderms that lived from the Early to Middle Devonian. Fossils appear in various strata in Russia, Central Asia, Australia, and China.
Arenipiscis westolli is an extinct buchanosteid arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata of New South Wales, Australia.
A plate in animal anatomy may refer to several things:
Asterolepis is an extinct genus of antiarch placoderms from the Devonian of North and South America and Europe. They were heavily armored flat-headed benthic detritivores with distinctive jointed limb-like pectoral fins and hollow spine. The armor plate gives the Asterolepis a box-like shape. Its pectoral fins are also armored but the caudal and dorsal fin are not. The first fossils were named by M. Eichwald in 1840 after noticing star-like markings on the fossils.
Osteoderms are dermal bone structures that support the upper layer of skin and serve as protection against the elements in a large variety of extinct and extant organisms, especially reptiles. This structure is commonly called "dermal armor" and serves to protect the organism, while also helping with temperature regulation. Osteoderms represent hard tissue components of the integument, making them easy to identify in fossil examination. This dermal armor is found prominently in many lizards. Some early amphibians have this armor, but it is lost in modern species with the exception a ventral plate, called the gastralia.
Qilinyu is an extinct genus of "maxillate" placoderm from the late Silurian of China. It contains a single species, Qilinyu rostrata, from the Xiaoxiang fauna of the Kuanti Formation. Along with its contemporary Entelognathus, Qilinyu is an unusual placoderm showing some traits more similar to bony fish, such as dermal jaw bones and lobe-like fins. It can be characterized by adaptations for a benthic lifestyle, with the mouth and nostrils on the underside of the head, similar to the unrelated antiarch placoderms. The shape of the skull has been described as "dolphin-like", with a domed cranium and a short projecting rostrum.
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