| Ctenurella | |
|---|---|
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| fossil of C. gladbachensis | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | † Placodermi |
| Order: | † Ptyctodontida |
| Family: | † Ptyctodontidae |
| Genus: | † Ctenurella Ørvig, 1960 |
Ctenurella ("Little Comb Tail") is an extinct genus of ptyctodont placoderm from the Late Devonian of Germany. The first fossils were found in the Strundevalley in the Paffrather Kalkmulde.
As with other ptyctodonts, the armor of Ctenurella was reduced to a few thin plates on the head and shoulder region. It was also relatively small for a placoderm, at just 13 centimetres (5 in) in length. It had two dorsal fins, with that at the rear of the body being relatively long and low, and large pectoral and pelvic fins. Most ptyctodonts are presumed to have fed on the ocean floor, but the well-developed fins of this genus indicate that it probably also swam in open waters. [1]
Ctenurella had a long, whip-like tail, large eyes, and robust upper and lower jaw tooth plates. Males also had hook-shaped sex organs, known as claspers. Since analogous features are also found in the unrelated living chimaeras, chimaeras and ptyctodonts are thought to be an example of convergent evolution. [1]
The specific name of the type species, C. gladbachensis, is a reference of the place Bergisch Gladbach, where it was founded. [2]
The species Ctenurella gardineri was split off into the new genus Austroptyctodus by Long (1997) in his review of the Gogo ptyctodontid species, stipulating that the genus Ctenurella as only coming from the German sites. A new description of Ctenurella gladbachensis by Long (1997) showed that the original restoration had wrongly restored the skull-roof as the central bones do not in fact meet each other behind the nuchal plate.
Ptyctodontid placoderms recently have been shown to give birth to live young, with specimens of pregnant females from two genera, Materpiscis and Austroptyctodus , both from the Gogo Formation of Western Australia, showing the presence of unborn embryos within the mother fishes (Long et al. 2008)
Placodermi is a class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scaled or naked, depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyletic, consisting of several distinct outgroups or sister taxa to all living jawed vertebrates, which originated among their ranks. This is illustrated by a 419-million-year-old fossil, Entelognathus, from China, which is the only known placoderm with a type of bony jaw like that found in modern bony fishes. This includes a dentary bone, which is found in humans and other tetrapods. The jaws in other placoderms were simplified and consisted of a single bone. Placoderms were also the first fish to develop pelvic fins, the precursor to hindlimbs in tetrapods, as well as true teeth. Paraphyletic groupings are problematic, as one can not talk precisely about their phylogenic relationships, characteristic traits, and complete extinction. 380-million-year-old fossils of three other genera, Incisoscutum, Materpiscis and Austroptyctodus, represent the oldest known examples of live birth. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic.
Dunkleosteus is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 358–382 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: D. terrelli, D. belgicus, D. denisoni, D. marsaisi, D. magnificus, D. missouriensis, D. newberryi, D. amblyodoratus, and D. raveri. The largest and most well known species is D. terrelli, which grew up to 8.79 m (28.8 ft) long and 4 t in weight. Dunkleosteus could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of 6,000 N at the tip and 7,400 N at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco.
Bothriolepis was a widespread, abundant and diverse genus of antiarch placoderms that lived during the Middle to Late Devonian period of the Paleozoic Era. Historically, Bothriolepis resided in an array of paleo-environments spread across every paleocontinent, including near shore marine and freshwater settings. Most species of Bothriolepis were characterized as relatively small, benthic, freshwater detritivores, averaging around 30 centimetres (12 in) in length. However, the largest species, B. rex, had an estimated bodylength of 170 centimetres (67 in). Although expansive with over 60 species found worldwide, comparatively Bothriolepis is not unusually more diverse than most modern bottom dwelling species around today.
Ischyodus is an extinct genus of cartilaginous fish belonging to the subclass Holocephali, which includes the modern-day chimaeras. Fossils are known from Europe, North America, and New Zealand.
Groenlandaspis is an extinct genus of arthrodire from the Late Devonian. Fossils of the different species are found in late Devonian strata in all continents except eastern Asia. The generic name commemorates the fact that the first specimens of the type species (G.mirabilis) were found in Greenland.
Acanthothoraci is an extinct group of chimaera-like placoderms who were closely related to the rhenanid placoderms. Superficially, the acanthoracids resembled scaly chimaeras, or (relatively) heavily armored ptyctodonts. They were distinguished from chimaeras by the presence of large scales and plates, a pair of large spines that emanate from their chests, tooth-like beak plates, and the typical bone-enhanced placoderm eyeball. They were distinguished from other placoderms due to differences in the anatomy of their skulls, and due to patterns on the skull plates and thoracic plates that are unique to this order.
The ptyctodontids ("folded-teeth") are placoderms of the order Ptyctodontida, containing the family Ptyctodontidae. With their big heads, big eyes, reduced armor and long bodies, the ptyctodontids bore a superficial resemblance to modern day chimaeras (Holocephali). Their armor was reduced to a pattern of small plates around the head and neck. Like the extinct and related acanthothoracids, and the living and unrelated holocephalians, most of the ptyctodontids are thought to have lived near the sea bottom and preyed on shellfish.
Campbellodus decipiens is an extinct ptyctodontid placoderm fish that lived around 380 million years ago. Its fossil remains have been found preserved in perfect three-dimensional form from the Gogo Formation of Western Australia. Originally it was described from large tooth plates and isolated skull roof bones by Miles & Young (1977). Long (1995) restored the complete fish based on new material found at Gogo in the mid 1980s, and described by Long (1997).
Onychodus is a genus of prehistoric lobe-finned fish which lived during the Devonian period. It is one of the best known of the group of onychodontiform fishes. Scattered fossil bones of Onychodus were first discovered in 1857, in North America, and described by John Strong Newberry. Other species were found in Australia, England, Norway and Germany showing that it had a widespread range.
Phyllolepis is the type genus of Phyllolepida, an extinct taxon of arthrodire placoderm fish from the middle to late Devonian. The species of Phyllolepis, themselves, are restricted to the Famennian-aged freshwater strata of the Late Devonian, around 360 million years ago. Fossils of this genus have been found primarily in Europe and North America. The end of the Devonian saw them disappear in a mass extinction.
Materpiscis is a genus of ptyctodontid placoderm from the Late Devonian located at the Gogo Formation of Western Australia. Known from only one specimen, it is unique in having an unborn embryo present inside the mother, with remarkable preservation of a mineralised placental feeding structure. This makes Materpiscis the oldest known vertebrate to show viviparity, or giving birth to live young.
Eastmanosteus is a fossil genus of dunkleosteid placoderms. It was closely related to the giant Dunkleosteus, but differed from that genus in size, in possessing a distinctive tuberculated bone ornament, a differently shaped nuchal plate and a more zig-zagging course of the sutures of the skull roof.
Incisoscutum is a genus of arthrodire placoderm from the Late Frasnian Gogo Reef, from Late Devonian Australia. The genus contains two species I. ritchiei, named after Dr. Alex Ritchie, a palaeoichthyologist and senior fellow of the Australian Museum, and I. sarahae, named after Sarah Long, daughter of its discoverer and describer, Dr. John A. Long.
Deinodus is a form genus that includes two species: the form found in the Onondaga Formation of western New York, Deinodus bennetti, and the form found in the Columbus and Limestone of central Ohio, Deinodus ohioensis. Both species are limited to the Eifelian age of the middle Devonian Period, which occurred 398-391 million years ago.
Austroptyctodus gardineri is a small ptyctodontid placoderm fish from the Upper Devonian Gogo Formation of Western Australia. First described by Miles & Young (1977) as a new species of the German genus Ctenurella. Long (1997) redescribed the German material and found major differences in the skull roof pattern so assigned it to a new genus, Austroptyctodus. This genus lacks spinal plates and has Ptyctodus-like toothplates.
Pinguosteus thulborni is a species of arthrodire placoderm from the Gogo Reef Formation, of Late Devonian Australia. The proportions of its armor, coupled with the relative lack of otherwise diagnostic ornamentation on the armor make its classification difficult, and so it was placed as incertae sedis within the placoderm group Coccosteina. It is the only known species of the genus Pinguosteus.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Microbrachius is an extinct genus of tiny, advanced antiarch placoderms closely related to the bothriolepids. Complete articulated specimens show that the armored section of the body had an average length of 2-4 cm. Species of Microbrachius are characterized by having large heads with short thoracic armor. Specimens of Microbrachius have been found in Scotland, Belarus, Estonia, and China. Specimens range in age from the Lower Devonian Late Emsian Stage to the Middle Devonian Upper Givetian Stage.
Asterolepis is an extinct genus of antiarch placoderms from the Devonian of North and South America and Europe. They were heavily armored flat-headed benthic detritivores with distinctive jointed limb-like pectoral fins and hollow spine. The armor plate gives the Asterolepis a box-like shape. Its pectoral fins are also armored but the caudal and dorsal fin are not. The first fossils were named by M. Eichwald in 1840 after noticing star-like markings on the fossils.
Ptyctodus is an extinct armour-plated fish of the late Devonian. Ptyctodus belongs to the family Ptyctodontidae and is of the class Placodermi. They share a close resemblance to modern day chimaeras (Holocephali). Fossils of this armour-plated fish have been found in locations such as in Russia, the Michigan Basis, and Arizona, United States.
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