Jackal | |
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Golden jackal (Canis aureus) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Canidae |
Subfamily: | Caninae |
Tribe: | Canini |
Subtribe: | Canina |
Species referred to as jackals | |
Jackals are canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed jackal (Lupulella mesomelas) and side-striped jackal (Lupulella adusta) of Central and Southern Africa, and the golden jackal (Canis aureus) of south-central Europe and Asia. The African golden wolf (Canis lupaster) was also formerly considered a jackal.
While they do not form a monophyletic clade, all jackals are opportunistic omnivores, predators of small to medium-sized animals and proficient scavengers. Their long legs and curved canine teeth are adapted for hunting small mammals, birds, and reptiles, and their large feet and fused leg bones give them a physique well-suited for long-distance running, capable of maintaining speeds of 16 km/h (10 mph) for extended periods of time. Jackals are crepuscular, most active at dawn and dusk.
Their most common social unit is a monogamous pair, which defends its territory from other pairs by vigorously chasing intruders and marking landmarks around the territory with their urine and feces. The territory may be large enough to hold some young adults, which stay with their parents until they establish their own territories. Jackals may occasionally assemble in small packs, for example, to scavenge a carcass, but they normally hunt either alone or in pairs.
The English word "jackal" dates back to 1600 and derives from the French chacal, itself from Turkish çakal, itself from Persian شغالshaghāl, from or cognate with Sanskrit शृगाल śṛgāla- meaning "the howler". [1] [2]
The extant wolf-like canids |
Phylogenetic relationships between the extant wolf-like clade of canids based on mitochondrial DNA. [3] [4] |
Similarities between jackals and coyotes led Lorenz Oken, in the third volume of his Lehrbuch der Naturgeschichte (1815), to place these species into a new separate genus, Thos, named after the classical Greek word θώς "jackal", but his theory had little immediate impact on taxonomy at the time. Angel Cabrera, in his 1932 monograph on the mammals of Morocco, questioned whether or not the presence of a cingulum on the upper molars of the jackals and its corresponding absence in the rest of Canis could justify a subdivision of that genus. In practice, Cabrera chose the undivided-genus alternative and referred to the jackals as Canis instead of Thos. [5]
Oken's Thos theory was revived in 1914 by Edmund Heller, who embraced the separate genus theory. Heller's names and the designations he gave to various jackal species and subspecies live on in current taxonomy, although the genus has been changed from Thos to Canis. [5]
The wolf-like canids are a group of large carnivores that are genetically closely related. They all have 78 chromosomes. The group includes genus Canis, Cuon, and Lycaon. The members are the dog (C. lupus familiaris), gray wolf (C. lupus), coyote (C. latrans), golden jackal (C. aureus), Ethiopian wolf (C. simensis), black-backed jackal (C. mesomelas), side-striped jackal (C. adustus), dhole (Cuon alpinus), and African wild dog (Lycaon pictus). [6] The latest recognized member is the African wolf (C. lupaster), which was once thought to be an African branch of the golden jackal. [4] As they possess 78 chromosomes, all members of the genus Canis are karyologically indistinguishable from each other, and from the dhole and the African hunting dog. [7] [8] The two African jackals are shown to be the most basal members of this clade, indicating the clade's origin from Africa. [3] Canis arnensis arrived in Mediterranean Europe 1.9 million years ago and is probably the ancestor of modern jackals. [9]
The paraphyletic nature of Canis with respect to Lycaon and Cuon has led to suggestions that the two African jackals should be assigned to different genera, Schaeffia for the side-striped jackal and Lupulella for the black-backed jackal [10] or Lupulella for both. [11] [12]
The intermediate size and shape of the Ethiopian wolf has at times led it to be regarded as a jackal, thus it has also been called the "red jackal" or the "Simien jackal".
Species | Binomial authority | Description | Range |
---|---|---|---|
Black-backed jackal Lupulella mesomelas | Schreber, 1775 | The most lightly built jackal, once considered to be the oldest living member of the genus Canis, [13] it is now placed in the genus Lupulella. It is the most aggressive of the jackals, being known to attack animal prey many times its own weight, and it has more quarrelsome intrapack relationships. [14] | Southern Africa and eastern coast of Kenya, Somalia, and Ethiopia |
Side-striped jackal Lupulella adustus | Sundevall, 1847 | It primarily resides in wooded areas, unlike other jackal species. It is the least aggressive of the jackals, rarely preying on large mammals. [15] | Central and southern Africa |
Golden jackal Canis aureus | Linnaeus, 1758 | The largest and most widespread of the jackals, it is more closely related to wolves than to African jackal species. | Southeastern Europe, Middle East, western Asia, and South Asia |
Like foxes and coyotes, jackals are often depicted as clever sorcerers in the myths and legends of their regions. They are mentioned roughly 14 times in the Bible. It is frequently used as a literary device to illustrate desolation, loneliness, and abandonment, with reference to its habit of living in the ruins of former cities and other areas abandoned by humans. It is called "wild dog" in several translations of the Bible. In the King James Bible, Isaiah 13:21 refers to 'doleful creatures', which some commentators suggest are either jackals or hyenas. [16]
In the Indian Panchatantra stories, the jackal is mentioned as wily and wise. [17] In Bengali tantrik tradition, they represent the goddess Kali. It is said she appears as jackals when meat is offered to her.
The Serer religion and creation myth posits jackals were among the first animals created by Roog, the supreme deity of the Serer people. [18]
Canidae is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid. The family includes three subfamilies: the Caninae, and the extinct Borophaginae and Hesperocyoninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other species.
The dhole is a canid native to South, East and Southeast Asia. It is anatomically distinguished from members of the genus Canis in several aspects: its skull is convex rather than concave in profile, it lacks a third lower molar and the upper molars possess only a single cusp as opposed to between two and four. During the Pleistocene, the dhole ranged throughout Asia, with its range also extending into Europe but became restricted to its historical range 12,000–18,000 years ago. It is now extinct in Central Asia, parts of Southeast Asia, and possibly the Korean peninsula and Russia.
Canis is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails.
The Ethiopian wolf, also called the red jackal, the Simien jackal or Simien fox, is a canine native to the Ethiopian Highlands. In southeastern Ethiopia, it is also known as the horse jackal. It is similar to the coyote in size and build, and is distinguished by its long and narrow skull, and its red and white fur. Unlike most large canids, which are widespread, generalist feeders, the Ethiopian wolf is a highly specialised feeder of Afroalpine rodents with very specific habitat requirements. It is one of the world's rarest canids, and Africa's most endangered carnivore.
The black-backed jackal, also called the silver-backed jackal, is a medium-sized canine native to eastern and southern Africa. These regions are separated by roughly 900 kilometers.
The golden jackal, also called the common jackal, is a wolf-like canid that is native to Eurasia. The golden jackal's coat varies in color from a pale creamy yellow in summer to a dark tawny beige in winter. It is smaller and has shorter legs, a shorter tail, a more elongated torso, a less-prominent forehead, and a narrower and more pointed muzzle than the Arabian wolf. It is listed as Least Concern on the IUCN Red List due to its widespread distribution and high density in areas with plenty of available food and optimum shelter.
The side-striped jackal is a canine native to Central and Southern Africa.
Canid hybrids are the result of interbreeding between the species of the subfamily Caninae.
The Egyptian wolf or the Egyptian jackal is a subspecies of African golden wolf native to northern, eastern and a part of western Africa.
Caninae is the only living subfamily within Canidae, alongside the extinct Borophaginae and Hesperocyoninae. They first appeared in North America, during the Oligocene around 35 million years ago, subsequently spreading to Asia and elsewhere in the Old World at the end of the Miocene, some 7 million to 8 million years ago.
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
The African wild dog, also known as the painted dog or Cape hunting dog, is a wild canine native to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus Lycaon, which is distinguished from Canis by dentition highly specialised for a hypercarnivorous diet and by a lack of dewclaws.
The African wolf is a canine native to North Africa, West Africa, the Sahel, northern East Africa, and the Horn of Africa. It is listed as least concern on the IUCN Red List. In the Middle Atlas in Morocco, it was sighted in elevations as high as 1,800 m (5,900 ft). It is primarily a predator of invertebrates and mammals as large as gazelle fawns, though larger animals are sometimes taken. Its diet also includes animal carcasses, human refuse, and fruit. They are monogamous and territorial; offspring remain with the parents to assist in raising their parents' younger pups.
It is widely agreed that the evolutionary lineage of the grey wolf can be traced back 2 million years to the Early Pleistocene species Canis etruscus, and its successor the Middle Pleistocene Canis mosbachensis. The grey wolf Canis lupus is a highly adaptable species that is able to exist in a range of environments and which possesses a wide distribution across the Holarctic. Studies of modern grey wolves have identified distinct sub-populations that live in close proximity to each other. This variation in sub-populations is closely linked to differences in habitat – precipitation, temperature, vegetation, and prey specialization – which affect cranio-dental plasticity.
Canis arnensis, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. Canis arnensis has been described as a small jackal-like canid. Its anatomy and morphology relate it more to the modern golden jackal than to the larger Etruscan wolf of that time. It is probably the ancestor of modern jackals.
Canis etruscus, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. The Etruscan wolf is described as a small wolf-like dog. It is widely agreed to be the ancestor of Canis mosbachensis, and thus ultimately the modern grey wolf.
Canis mosbachensis is an extinct wolf that inhabited Europe from the late Early Pleistocene to the Middle Pleistocene, around 1.4 million to 400,000 years ago. Canis mosbachensis is widely considered to have descended from the earlier Canis etruscus, and to be the ancestor of the living grey wolf with some considering it as a subspecies of the wolf as Canis lupus mosbachensis. The morphological distinction between C. mosbachensis and C. lupus has historically been vague, and attribution of fossils to C. mosbachensis or to C. lupus around the transition time between the two species is ambiguous.
This is a list of the species of Canidae ordered by average weights of adult individuals in the wild. It does not include canid hybrids or any domesticated animals. Only wild species of canids are included, all of which are described as species by authentic sources.
Lupulella is a genus of canine found in Africa. This genus consists of only two extant species, the black-backed jackal and the side-striped jackal.