Macroloxoceras

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Macroloxoceras
Temporal range: Upper Devonian
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Macroloxoceras

Macroloxoceras is a large pseuorthocerid from the upper Devonian of Central Colorado and Southern New Mexico with features resembling those found in actinocerids. [1] Pseudorthocerids and actinocerids (respectively Pseuorthocerida and Actinocerida) are extinct nautiloid cephalopods, generally with long straight shells and expanded siphuncle segments filled with organic deposits.

Pseudorthocerida is an order of generally straight longiconic Orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining.. Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature

The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian, 419.2 million years ago (Mya), to the beginning of the Carboniferous, 358.9 Mya. It is named after Devon, England, where rocks from this period were first studied.

Colorado State of the United States of America

Colorado is a state of the Western United States encompassing most of the southern Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains. It is the 8th most extensive and 21st most populous U.S. state. The estimated population of Colorado was 5,695,564 on July 1, 2018, an increase of 13.25% since the 2010 United States Census.

Contents

Morphologic Description

Macroloxoceras has an orthconic shell with a strongly depressed cross section and markedly flattened venter. Sutures have broad ventral lobes but are otherwise straight and transverse. The siphuncle is ventral of the center; composed of broadly expanded segments with a spheroidal outline. Septal necks are cytochoantic. Connecting rings are thin and make contact over a wide area on the previous septa at their adapical ends. Endosiphuncular deposits are fairly ubiquitous, begin at the septal openings and grow mostly forward to connect with the next in the anterior part of the following segment. Cavities left in the segments are connected to the rings by two series of radial canals; one at the anterior end of the segment near the end of the septal neck; the other starting in the middle, curving back and ending about where the connecting rings makes contact with the previous septum. Cameral deposits are well developed. [1] [2]

In anatomy, a suture is a fairly rigid joint between two or more hard elements of an organism, with or without significant overlap of the elements.

Taxonomy

Macroloxoceras, named and described by Rousseau H. Flower in 1957, is included in the Pseudorthocerid family Pseudactinoceratidae and is placed in the subfamily Macroloxoceratinae, named by him for this genus. [1] Flower in 1957 designated the Pseuorthoceratidae as a family in the Michelinoceratida, more commonly known as the Orthocerida. [2]

Rousseau Hayner Flower (1913–1988) was an extremely prolific 20th century paleontologist, known for his eccentric personality.

Two species have been identified, the genotype Macroloxoceras magnum from the Upper Devonian Chaffe Limestone near Glenwood Springs, Colorado and Macroloxoceras minor from the upper 20 ft of the Pecha Shale near Santa Rita, New Mexico. [1]

The holotype of M. magnum is a 240 millimetres (9.4 in) long section of the phragmocone with 14 camerae and s short portion of the living chamber, the cross section of which increases from a width of 56 millimetres (2.2 in) and height of 37mm at the base to a width of 100 millimetres (3.9 in) and height of about 42 millimetres (1.7 in) at the anterior end. The chambers increase in length from 10–19 millimetres (0.39–0.75 in). Septal openings are 5mm across, rings expand so as segments are at least 16 millimetres (0.63 in) at their greatest width. Siphunclular deposits are thickest in the posterior part of the segments, become thin in the anterior part where they connect with the next deposit forward. Radial canals are as described for the genus. [1]

The holotype of M. minor is a portion of a phragmocone with a natural horizontal section of the siphuncle with the ventral portion remaining and dorsal portion removed be erosion.

The M. magnum holotype is deposited with the Paleontological Research Institute while the M. minor holotype is in the paleontology collection of the New Mexico Museum of Natural History and Science, transferred from the New Mexico Bureau of Mines and Mineral Resources.

Related Research Articles

Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician.

Lambeoceras is a genus of rather large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae.

Paraloxoceras is a genus of straight shelled, orthoconic nautiloid cephalopods, now extinct, that lived during the Early Carboniferous. Fossils have been found in Europe and central Asia; the type, P. konincki, named by Flower, came from Belgium.

Spyroceras is a genus of pseudorthocerids from the Devonian of North America and Europe, defined by Hyatt in 1884. Pseudorthocerids are a kind of orthocertaoid, a taxonomic group within the Nautiloidea. Specifically Spyroceras belongs to the pseudorthocerid family, Spyroceratidae.

Pseudactinoceras is an extinct nautiloid cephalopod included in the order Pseudorthocerida and the namesake of the family Pseudactinoceratidae.

Wardoceras is an extinct nautiloid genus from the late Early Ordovician of Western Utah, assigned to the orthocerid family, Michelinoceratidae

Metabaltoceras, first described by Rousseau Flower in 1964, is a fossil cephalopod genus in the family Baltoceratidae, with a small, slender, fairly fusiform shell, and a large empty siphuncle in contact with the ventral surface. The siphuncle is achoanitic, having virtually no septal necks, segments formed all but entirely by connecting rings. Shells are generally straight, beginning with a subcircular cross section in the adapical portion but becoming faintly depressed adorally. Sutures are straight and transverse except ventrally, where they produce deep, prominent lobes.

Sactoceras is an extinct nautiloid cephalopod that lived during the Ordovician and Silurian in what would become North America, Europe, and Asia.

Carbactinoceratidae is a family of extinct cephalopods with external shells that lived around 325 million years ago, during the Carboniferous period. They were the last group of orthocones to attain sizes exceeding one meter in length. One specimen of the carbactinoceratid species Rayonnoceras solidiforme, recently found in Arkansas, was measured at 2.5 metres (8.2 ft) long.

The Actinoceriatidae are a family of actinocerids named by Saemann in 1853 for those that grew to have large shells with blunt apecies and large siphuncles with widely expanded segments and a generally arcuate endosiphucular canal system. Their range is from the upper Middle Ordovician to the Lower Silurian. Actinocerids are generally straight-shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals

<i>Armenoceratidae</i> family of molluscs

The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida.

Nybyoceras is an actinocerid genus assigned to the Armenoceratidae and similar to Armenoceras except for having a siphuncle close to the ventral side of the shell.

Pseudactinoceraidae is a family in the nautiloid cephalopod order, Pseudorthocerida, known from the Upper Devonian and Lower Carboniferous (Mississippian).

Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber. Apertures vary from round to T-shaped, with a long narrow, ventral, hyponomic sinus for the maneuvering funnel. Siphuncles are commonly central or subcentral with generally broad, expanded, segments and usually thin connecting rings.

Veneficoceras is a genus of the rod-bearing Baltoceratidae, an extinct cephalopod family with characteristics of the orthoceratoid Dissidocerida, found in Cassinianage, Lower Ordovician, limestone in western Utah.

Kobyashiceras is an extinct genus in the cephalopod order Actinocerida, from Lower Devonian marine sediments in Japan. The type and sole included species is Kobayashiceras gifuense. The generic name honors the late Dr. Teiichi Kobayashi who contributed greatly to the study of Paleozoic cephalopods. The specific name is derived from Gifu, the name of the prefecture in which the type locality is found.

The Sactorthoceratidae comprise Orthocerataceaen genera with a subcentral suborthochoanitic siphuncle composed of slightly expanded segments and free of organic deposits. The camerae (chambers) of the phragmocone likewise have organic deposits that are typically retarded or sparse.

Adnatoceras is a genus of mid Devonian to Pennsylvanian orthoconic nautiloid cephalopods included in the pseudorthocerid family Spyroceratidae, characterized by tubular, straight-sided, siphuncle segments that expand abruptly at the septal foremena. In some species the siphuncle segments become more ovoid in the later grow stages. The name Adnatoceas comes from the wide area of contact, or adnation, between the connecting ring and flared out brim of the cyrtochaoanitic septal neck.

Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.

Dolorthoceras is a nautiloid cephalopod from the upper Paleozoic found in Lower Devonian to Lower Permian strata in North America, Europe, Asia, and Australia.

References

  1. 1 2 3 4 5 Macroloxoceras, a Devonian Homeomorph of the Actinoceratida, part II Memoir 2, Studies of the Actinoceatidae by Rousseau H Flower. New Mexico Bureau of Mines and Mineral Resources, Socorro NM 1957
  2. 1 2 Nautiloidea-Orthocerida by Walter C Sweet, Treatise on Invertebrate Paleontology Part K Mollusca 3, R.C Moore (ed) 1964