Manis palaeojavanica Temporal range: Pleistocene late | |
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bones of Manis palaeojavanica (darker) compared to same bones of Manis javanica (lighter) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Pholidota |
Family: | Manidae |
Genus: | Manis |
Subgenus: | Paramanis |
Species: | †M. palaeojavanica |
Binomial name | |
†Manis palaeojavanica Dubois, 1907 [1] | |
Synonyms | |
Giant asian pangolin (Manis palaeojavanica ["ancient Javan pangolin"]), is an extinct species of pangolin (from genus Manis ) that was native to Asia. [5]
In 1926, E. Dubois described the bones of M. palaeojavanica discovered in Java. Later, Lord Medway excavated another set of bones at the Niah Caves in Malaysia. In 1960, D. A. Hoojier identified these bones as that of an extinct species. Using carbon dating, the Niah Caves bones were determined to be 42,000–47,000 years old. [6]
M. palaeojavanica is one of the only known fossils to be found nearly complete of the pangolin species, due to their armour-like scales that protected their bodies when they were alive. [7] Its total length is measured up to 2.5 m (8.2 ft). [8]
Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota. The one extant family, the Manidae, has three genera: Manis, Phataginus, and Smutsia. Manis comprises four species found in Asia, while Phataginus and Smutsia include two species each, all found in sub-Saharan Africa. These species range in size from 30 to 100 cm. A number of extinct pangolin species are also known. In September 2023, nine species were reported.
Solo Man is a subspecies of H. erectus that lived along the Solo River in Java, Indonesia, about 117,000 to 108,000 years ago in the Late Pleistocene. This population is the last known record of the species. It is known from 14 skullcaps, two tibiae, and a piece of the pelvis excavated near the village of Ngandong, and possibly three skulls from Sambungmacan and a skull from Ngawi depending on classification. The Ngandong site was first excavated from 1931 to 1933 under the direction of Willem Frederik Florus Oppenoorth, Carel ter Haar, and Gustav Heinrich Ralph von Koenigswald, but further study was set back by the Great Depression, World War II and the Indonesian War of Independence. In accordance with historical race concepts, Indonesian H. erectus subspecies were originally classified as the direct ancestors of Aboriginal Australians, but Solo Man is now thought to have no living descendants because the remains far predate modern human immigration into the area, which began roughly 55,000 to 50,000 years ago.
Sundaland is a biogeographical region of South-eastern Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It includes Bali, Borneo, Java, and Sumatra in Indonesia, and their surrounding small islands, as well as the Malay Peninsula on the Asian mainland.
Java Man is an early human fossil discovered in 1891 and 1892 on the island of Java. Estimated to be between 700,000 and 1,490,000 years old, it was, at the time of its discovery, the oldest hominid fossil ever found, and it remains the type specimen for Homo erectus.
Manis ("spirit") is a genus of South Asian and East Asian pangolins, the Asiatic pangolins, from subfamily Maninae, within family Manidae.
Manidae ("spirits") is the only extant family of pangolins from superfamily Manoidea. This family comprises three genera, as well as extinct Fayum pangolin.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition. Some modern populations of Asian elephants have also undergone size reduction on islands to a lesser degree, resulting in populations of pygmy elephants.
The Sunda pangolin, also known as the Malayan or Javan pangolin, is a species of pangolin.
Trinil is a palaeoanthropological site on the banks of the Bengawan Solo River in Ngawi Regency, East Java Province, Indonesia. It was at this site in 1891 that the Dutch anatomist Eugène Dubois discovered the first early hominin remains to be found outside of Europe: the famous "Java Man" specimen.
Panthera tigris trinilensis, known as the Trinil tiger, is an extinct tiger subspecies. The Trinil tiger is known from remains dating from nearly 1.2 million years ago, which were found at the locality of Trinil, Java, Indonesia. The fossil remains are now stored in the Dubois Collection of the National Museum of Natural History in Leiden, Netherlands. Although these fossils have been found on Java, the Trinil tiger is thought not to be a direct ancestor of the Javan tiger, and likely went extinct ca. 50,000 years ago. Additionally, the Bali tiger, also extinct, was also not closely related to the Trinil tiger due to the different time periods in which they were alive.
The lesser false vampire bat is a bat found in South Asia and Southeast Asia from Sri Lanka and India in the west to Indonesia and the Philippines in the east. They live in caves and tree hollows. They are insectivorous.
The Chinese pangolin is a pangolin native to the northern Indian subcontinent, northern parts of Southeast Asia and southern China. It has been listed as Critically Endangered on the IUCN Red List since 2014, as the wild population is estimated to have declined by more than 80% in three pangolin generations, equal to 21 years. It is threatened by poaching for the illegal wildlife trade.
The Palawan bearded pig is a pig species in the genus Sus endemic to the Philippines, where it occurs on the archipelago of islands formed by Balabac, Palawan, and the Calamian Islands. It is 1 to 1.6 m in length, about 1 m (3.3 ft) tall and weigh up to 150 kg (330 lb).
Bos palaesondaicus occurred on Pleistocene Java (Indonesia) and belongs to the Bovinae subfamily. It has been described by the Dutch paleoanthropologist Eugène Dubois in 1908. The holotype of Bos palaesondaicus is a skull from Trinil. This species is the likely ancestor to the banteng.
The island of Borneo is located on the Sunda Shelf, which is an extensive region in Southeast Asia of immense importance in terms of biodiversity, biogeography and phylogeography of fauna and flora that had attracted Alfred Russel Wallace and other biologists from all over the world.
The region of Southeast Asia is considered a possible place for the evidence of archaic human remains that could be found due to the pathway between Australia and mainland Southeast Asia, where the migration of multiple early humans has occurred out of Africa. One of many pieces of evidence is of the early human found in central Java of Indonesia in the late 19th century by Eugene Dubois, and later in 1937 at Sangiran site by G.H.R. van Koenigswald. These skull and fossil materials are Homo erectus, named Pithecanthropus erectus by Dubois and Meganthropus palaeojavanicus by van Koenigswald. They were dated to c. 1.88 and 1.66 Ma, as suggested by Swisher et al. by analysis of volcanic rocks.
Duboisia santeng or Dubois' antelope is an extinct antelope-like bovid that was endemic to Indonesia during the Pleistocene. It went extinct during the Ionian stage of the Pleistocene, about 750.000 years ago. Duboisia santeng was first described by the Dutch paleoanthropologist and geologist Eugène Dubois in 1891.
Mececyon trinilensis, the Trinil dog, is an extinct canid species that lived on the island of Java in Indonesia during the Pleistocene.
Gavialis bengawanicus is an extinct species of crocodilian that is related to the modern Indian gharial. Fossils have been found in Thailand and Indonesia. The type locality is at Trinil.
Smutsia olteniensis is an extinct species of pangolins from genus Smutsia of subfamily Smutsiinae within family Manidae. Fossilized remains of the species were found in Romania, providing evidence regarding the existence of pangolins in Europe during the Plio-Pleistocene period.