Odontoanserae | |
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The skull of Pelagornis mauretanicus (top) and the head of common merganser (bottom) showing the serrated edge common in this group | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Superorder: | Galloanserae |
Clade: | Odontoanserae Bourdon, 2005 |
Subgroups | |
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The Odontoanserae is a proposed clade that includes the family Pelagornithidae (pseudo-toothed birds) and the clade Anserimorphae (the order Anseriformes and their stem-relatives). [2] The placement of the pseudo-toothed birds in the evolutionary tree of birds has been problematic, with some supporting the placement of them near the orders Procellariformes and Pelecaniformes based on features in the sternum. [3]
In 2005 a cladistic analysis had found support in placing pseudo-toothed birds as the sister group to waterfowl. [2] Evidence for this comes from shared characteristics in the skull such as lack of a crest on the underside of the palatine bone and two condyles on the mandibular process of the quadrate bone, with the middle condyle beakwards of the side condyle. [2] In addition to that, both groups have similar features in their pelvic and pectoral regions. Furthermore, a 2013 study on the growth pattern and structure of the pseudoteeth in Pelagornis mauretanicus shows more support of Odontoanserae as both groups have "soft rhamphotheca, or delayed hardening of the rhamphotheca." [4] In addition to Pelagornithidae and Anseriformes paleontologists also have support in placing mihirungs (Dromornithidae) and Gastornithids into this group, as they too also share anatomical features in the skull and pelvic bones with waterfowl. [5] [6] [7] The mihirungs and the gastornithids are more derived than the pseudo-tooth birds and are closer to Anseriformes. One hypothesis is that diatrymas and mihirungs are successive sister groups to anseriforms and another hypothesis places mihirungs as crown anseriforms closely related to the screamers (Anhimidae). [6]
Below is the general consensus of the phylogeny. [6] [2] [7] [8] [9]
Odontoanserae |
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However, a 2017 paper by Worthy and colleagues found an alternative phylogeny concerning Anserimorphae. [10] By adding additional new characters, as well as incorporating several new taxa into established matrices, the authors have found gastornithids and mihirungs to be sister taxa which could be placed in the order Gastornithiformes. [10] In addition, they have found support that the family Vegaviidae (usually classified as crown anseriforms or their sister taxon [9] ) are more related to gastornithiforms than to anseriforms; accordingly, they have created the monotypic order Vegaviiformes. [10] The authors did note the bootstrap support is weakly supported and several alternative phylogenies in their paper found gastornithiforms to be stem galliforms instead. [10] These too were weakly supported as well. [10] Below is a simplified phylogeny showing their one phylogeny supporting gastornithiforms as anserimorphs. [10]
Anserimorphae |
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In 2019 a new species Conflicto antarcticus was described from Early Paleocene deposits in Antarctica. [1] Known completely from associated bones from a single individual, Tambussi et al. (2019) incorporated the new taxon into a phylogenetic analysis using the matrix data from Worthy et al. (2017). Their results not only supported the sister grouping of vegaviids with gastornithids and mihirungs (which included Vegaviidae into Gastornithiformes), but also found two taxa, Anatalavis rex and the tall, wading presbyornithids, traditionally placed as part of the anseriform crown, [11] to be stem-anseriforms. [1] Below is the Tambussi et al. (2019) phylogeny. [1]
Anserimorphae |
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Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae, Anseranatidae, and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves. Due to their aquatic nature, most species are web-footed.
Gastornis is an extinct genus of large flightless birds that lived during the mid-Paleocene to mid-Eocene epochs of the Paleogene period. Fossils have been found in Europe, Asia and North America, with the remains from North America originally assigned to the genus Diatryma.
Vegavis is a genus of extinct bird that lived during the Late Cretaceous of Antarctica, some 68 to 66 mya. Among modern birds, most studies show that Vegavis is most closely related to ducks and geese (Anatidae), but it is not considered to be a direct ancestor of them, although other studies question these results.
Dromornithidae, known as mihirungs and informally as thunder birds or demon ducks, were a clade of large, flightless Australian birds of the Oligocene through Pleistocene Epochs. All are now extinct. They were long classified in Struthioniformes, but are now usually classified as galloanseres. Dromornithids were part of the Australian megafauna. One species, Dromornis stirtoni, was 3 m tall. Only a single species, Genyornis newtoni survived into the Late Pleistocene. They are thought to have been herbivorous.
Brontornis is an extinct genus of giant bird that inhabited Argentina during the Early to Middle Miocene. Its taxonomic position is highly controversial, with authors alternatively considering it to be a cariamiform, typically a phorusrhacid or an anserimorph.
Presbyornithidae is an extinct group of birds with a global distribution. They had evolved by the late Cretaceous period and became extinct during the early Miocene. Initially, they were believed to present a mix of characters shown by waterbirds, shorebirds and flamingos and were used to argue for an evolutionary relationship between these groups, but they are now generally accepted to be waterfowl closely related to modern ducks, geese, and screamers.
The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family of large seabirds. Their fossil remains have been found all over the world in rocks dating between the Early Paleocene and the Pliocene-Pleistocene boundary.
Gastornithiformes were an extinct order of giant flightless fowl with fossils found in North America, Eurasia, possibly Australia. Members of Gastornithidae were long considered to be a part of the order Gruiformes. However, the traditional concept of Gruiformes has since been shown to be polyphyletic.
Paraphysornis is an extinct genus of giant flightless terror birds that inhabited Brazil during Late Oligocene or Early Miocene epochs. Although not the tallest phorusrhacid, Paraphysornis measured up to 1.4 metres tall at the hips and weighed around 180–240 kilograms (400–530 lb). It was also a notably robust bird, having short and robust tarsal bones not suited for pursuit hunting.
Odontopteryx is a genus of the extinct pseudotooth birds or pelagornithids. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Cyphornis is a genus of the prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Dasornis is a genus of prehistoric pseudotooth birds. These were probably close relatives of either pelicans and storks or waterfowl; they are placed in the order Odontopterygiformes to account for this uncertainty.
Gigantornis eaglesomei is a very large prehistoric bird described from a fragmentary specimen from the Eocene of Nigeria. It was originally described as a representative of the albatross family, Diomedeidae, but was later referred to the pseudotooth or bony-toothed bird family, Pelagornithidae. Gigantornis is one of the largest pseudotooth birds, with an inferred wingspan of about 6 m (20 ft) it is among the largest birds ever.
Palaeochenoides is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Pelagornis is a widespread genus of prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or waterfowl, and are placed here in the order Odontopterygiformes to account for this uncertainty.
Tympanonesiotes is a somewhat doubtfully valid genus of the prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Macrodontopteryx is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These animals were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Vegaviidae is an extinct family of ornithurines, often regarded as stem-anseriforms, which existed during the Late Cretaceous and possibly the Paleocene. Fossils attributed to the family have been found in Canada, Chile, New Zealand, and Antarctica.
Conflicto antarcticus is a species of stem waterfowl whose fossils were found in the early Paleocene López de Bertodano Formation of Antarctica, the only species of its genus and the family Conflictonidae. It is characterized by it slender body and long legs, yet possesses a duck-like bill which indicates the form of beak evolved early in Anseriformes.
Wunketru is an extinct genus of waterfowl from the Eocene Las Flores Formation of Chubut Province, Argentina. The genus contains a single species, W. howardae, known from a partial skeleton previously classified as a species of Telmabates.