Anseriformes Temporal range: Maastrichtian [1] Stem groups present since | |
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Magpie goose, Anseranas semipalmata | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Clade: | Anserimorphae |
Order: | Anseriformes Wagler, 1831 |
Subtaxa | |
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Range of the waterfowl and allies |
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves, the clade consisting of all other modern birds except the galliformes and paleognaths. Due to their aquatic nature, most species are web-footed.
Anseriformes are one of only two types of modern bird to be confirmed present during the Mesozoic alongside the other dinosaurs, and in fact were among the very few birds to survive their extinction, along with their cousins, the Galliformes. These two groups only occupied two ecological niches during the Mesozoic, living in water and on the ground, while the toothed Enantiornithes were the dominant birds that ruled the trees and air. The asteroid that ended the Mesozoic destroyed all trees as well as animals in the open, a condition that took centuries[ citation needed ] to recover from. The Anseriformes and Galliformes are thought to have survived in the cover of burrows and water, and not to have needed trees for food and reproduction. [2]
The earliest known stem anseriform is the presbyornithid Teviornis from the Nemegt Formation of Mongolia. [3] Some members apparently surviving the KT extinction event, including presbyornithids, thought to be the common ancestors of ducks, geese, swans, and screamers, the last group once thought to be Galliformes, but now genetically confirmed to be closely related to geese. The first known duck fossils start to appear about 34 million years ago.
Waterfowl are the best-known examples of sexually antagonistic genital coevolution in vertebrates, causing genital adaptations to coevolve in each sex to advance control over mating and fertilization. Sexually antagonistic coevolution (or SAC) occurs as a consequence of sexual conflict between males and females, resulting in coevolutionary process that reduce fit, or that functions to decrease ease of having sex. [4]
The Anseriformes and the Galliformes (pheasants, etc.) belong to a common group, the Galloanserae. They are the most primitive neognathous birds, and as such they should follow the Palaeognathae (ratites and tinamous) in bird classification systems. Several unusual extinct families of birds like the albatross-like pseudotooth birds and the giant flightless gastornithids and mihirungs have been found to be stem-anseriforms based on common features found in the skull region, beak physiology and pelvic region. [5] [6] [7] [8] [9] [10] The genus Vegavis for a while was found to be the earliest member of the anseriform crown group but a recent 2017 paper has found it to be just outside the crown group in the family Vegaviidae. [11]
Below is the general consensus of the phylogeny of anseriforms and their stem relatives. [5] [6] [7] [8] [9] [11]
Odontoanserae |
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Anatidae systematics, especially regarding placement of some "odd" genera in the dabbling ducks or shelducks, is not fully resolved. See the Anatidae article for more information, and for alternate taxonomic approaches. Anatidae is traditionally divided into subfamilies Anatinae and Anserinae. [12] The Anatinae consists of tribes Anatini, Aythyini, Mergini and Tadornini. The higher-order classification below follows a phylogenetic analysis performed by Mikko's Phylogeny Archive [13] [14] and John Boyd's website. [15]
Unassigned Anatidae:
In addition, a considerable number of mainly Late Cretaceous and Paleogene fossils have been described where it is uncertain whether or not they are anseriforms. This is because almost all orders of aquatic birds living today either originated or underwent a major radiation during that time, making it hard to decide whether some waterbird-like bone belongs into this family or is the product of parallel evolution in a different lineage due to adaptive pressures.
Living Anseriformes based on the work by John Boyd. [15]
Anseriformes classification |
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Studies of the mitochondrial DNA suggest the existence of four branches – Anseranatidae, Dendrocygninae, Anserinae and Anatinae – with Dendrocygninae being a subfamily within the family Anatidae and Anseranatidae representing an independent family. [18] The clade Somaterini has a single genus Somateria .
The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating on the water surface, and, in some cases, diving in at least shallow water. The family contains around 174 species in 43 genera.
The Oxyurini are a tribe of the duck subfamily of birds, the Anatinae. It has been subject of considerable debate about its validity and circumscription. Some taxonomic authorities place the group in its own subfamily, the Oxyurinae. Most of its members have long, stiff tail feathers which are erected when the bird is at rest, and relatively large, swollen bills. Though their relationships are still enigmatic, they appear to be closer to swans and true geese than to the typical ducks. The highest diversity is found in the warmer parts of the Americas, but at least one species occurs in a major part of the world.
The diving ducks, commonly called pochards or scaups, are a category of duck which feed by diving beneath the surface of the water. They are part of Anatidae, the diverse and very large family that includes ducks, geese, and swans.
The Anatinae are a subfamily of the family Anatidae. Its surviving members are the dabbling ducks, which feed mainly at the surface rather than by diving. The other members of the Anatinae are the extinct moa-nalo, a young but highly apomorphic lineage derived from the dabbling ducks.
The Tadorninae is the shelduck-sheldgoose subfamily of the Anatidae, the biological family that includes the ducks and most duck-like waterfowl such as the geese and swans.
The term perching ducks is used colloquially to mean any species of ducks distinguished by their readiness to perch high in trees.
The shelducks, most species of which are found in the genus Tadorna, are a group of large birds in the Tadorninae subfamily of the Anatidae, the biological family that includes the ducks and most duck-like waterfowl such as the geese and swans.
The magpie goose is the sole living representative species of the family Anseranatidae. This common waterbird is found in northern Australia and southern New Guinea. As the species is prone to wandering, especially when not breeding, it is sometimes recorded outside its core range. The species was once also widespread in southern Australia but disappeared from there largely due to the drainage of the wetlands where the birds once bred. Due to their importance to Aboriginal people as a seasonal food source, as subjects of recreational hunting, and as a tourist attraction, their expansive and stable presence in northern Australia has been "ensured [by] protective management".
Anseranatidae, the magpie-geese, is a biological family of waterbirds. The only living species, the magpie goose, is a resident breeder in northern Australia and in southern New Guinea.
Anas is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was split into four separate genera. The genus now contains 31 living species. The name Anas is the Latin for "duck".
Pygmy geese are a group of very small "perching ducks" in the genus Nettapus which breed in the Old World tropics. They are the smallest of all wildfowl. As the "perching ducks" are a paraphyletic group, they need to be placed elsewhere. The initially assumed relationship with the dabbling duck subfamily Anatinae has been questioned, and it appears they form a lineage in an ancient Gondwanan radiation of waterfowl, within which they are of unclear affinities. An undescribed fossil species from the late Hemphillian of Jalisco, central Mexico, has also been identified from the distal end of a tarsometatarsus. It is only record of the genus in the New World.
The whistling ducks or tree ducks are a subfamily, Dendrocygninae, of the duck, goose and swan family of birds, Anatidae. In other taxonomic schemes, they are considered a separate family, Dendrocygnidae. Some taxonomists list only one genus, Dendrocygna, which contains eight living species, and one undescribed extinct species from Aitutaki of the Cook Islands, but other taxonomists also list the white-backed duck under the subfamily.
Anser is a waterfowl genus that includes the grey geese and the white geese. It belongs to the true goose and swan subfamily of Anserinae under the family of Anatidae. The genus has a Holarctic distribution, with at least one species breeding in any open, wet habitats in the subarctic and cool temperate regions of the Northern Hemisphere in summer. Some also breed farther south, reaching into warm temperate regions. They mostly migrate south in winter, typically to regions in the temperate zone between the January 0 °C (32 °F) and 5 °C (41 °F) isotherms.
Presbyornis is an extinct genus of presbyornithid bird from North America during the Paleogene period, between the Late Paleocene and Early Eocene.
Presbyornithidae is an extinct group of birds found in North America, South America, East Asia, Australia and possibly North Africa. They had evolved by the Maastrichtian age of the Late Cretaceous and became extinct during the Aquitanian age of the Early Miocene. The family contains the oldest known neognath, Teviornis from the Nemegt Formation of Mongolia.
Anatalavis is genus of prehistoric birds related to ducks and geese, perhaps to the magpie-goose in particular. Alternatively, it might have been a more basal lineage of Anserimorphae distinct from the living waterfowl, similar or even related to the roughly contemporary Conflicto antarcticus from the Danian of Antarctica.
Bradley Curtis Livezey was an American ornithologist with scores of publications. His main research included the evolution of flightless birds, the systematics of birds, and the ecology and behaviour of steamer ducks.
The Odontoanserae is a proposed clade that includes the family Pelagornithidae and the clade Anserimorphae. The placement of the pseudo-toothed birds in the evolutionary tree of birds has been problematic, with some supporting the placement of them near the orders Procellariformes and Pelecaniformes based on features in the sternum.