Branta

Last updated

Branta
Temporal range:
Late Miocene-Holocene
Branta canadensis -Smythe Park, Toronto, Canada-8.jpg
A Canada goose (Branta canadensis) swimming at Smythe Park, Toronto, Ontario, Canada
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Anseriformes
Family: Anatidae
Subfamily: Anserinae
Genus: Branta
Scopoli, 1769
Type species
Anas bernicla (Brant goose)
Linnaeus, 1758
Species

Branta bernicla
Branta canadensis
Branta hutchinsii
Branta leucopsis
Branta ruficollis
Branta sandvicensis
and see text

Contents

Synonyms

Nesochen Salvadori, 1895

The black geese of the genus Branta are waterfowl belonging to the true geese and swans subfamily Anserinae. They occur in the northern coastal regions of the Palearctic and all over North America, migrating to more southerly coasts in winter, and as resident birds in the Hawaiian Islands. Alone in the Southern Hemisphere, a self-sustaining feral population derived from introduced Canada geese is also found in New Zealand.

The black geese derive their vernacular name for the prominent areas of black coloration found in all species. They can be distinguished from all other true geese by their legs and feet, which are black or very dark grey. Furthermore, they have black bills and large areas of black on the head and neck, with white (ochre in one species) markings that can be used to tell apart most species. [note 1] As with most geese, their undertail and uppertail coverts are white. They are also on average smaller than other geese, though some very large taxa are known, which rival the swan goose and the black-necked swan in size.

The Eurasian species of black geese have a more coastal distribution compared to the grey geese (genus Anser ) which share the same general area of occurrence, not being found far inland even in winter (except for occasional stray birds or individuals escaped from captivity). This does not hold true for the American and Pacific species, in whose ranges grey geese are, for the most part, absent.

Taxonomy

The genus Branta was introduced by the Austrian naturalist Giovanni Antonio Scopoli in 1769. [1] The name is a Latinised form of Old Norse Brandgás meaning burnt as in "burnt (black) goose". [2] The type species is the brant goose (Branta bernicla). [3]

Ottenburghs and colleagues published a study in 2016 that established the phylogenetic relationships between the species. [4]

Branta 

Brant (Branta bernicla)

Red-breasted goose (Branta ruficollis)

Nene (Branta sandvicensis)

Canada goose (Branta canadensis)

Barnacle goose (Branta leucopsis)

Cackling goose (Branta hutchinsii)

Species list

The genus contains six living species. [5]

ImageScientific nameCommon nameDistribution
Branta bernicla bernicla - Brent Goose.jpg Branta bernicla Brant (U.S.) or brent goose (U.K.)High Arctic tundra. Circumpolar; several distinct breeding populations, which winter in particular areas (some of which overlap) along the northern temperate zone of the Atlantic and Pacific coastlines. Some resident populations can be found [ citation needed ] in the United States and Western Europe.
Branta ruficollis 2.jpg Branta ruficollis Red-breasted goose Breeds in Arctic Siberia, mainly on the Taymyr Peninsula, with smaller populations in the Gydan and Yamal Peninsulas; wintering on northwestern shores of the Black Sea in Bulgaria, Romania and Ukraine, as well as in Azerbaijan
Branta sandvicensis LC399.jpg Branta sandvicensis Nene, nēnē, or Hawaiian gooseHawaiian Islands
Kanadagans Branta canadensis.jpg Branta canadensis Canada goose Temperate regions of North America, introduced populations in Western and northern Europe, Japan, New Zealand, Chile, Argentina and the Falkland Islands
Barnacle-Goose.jpg Branta leucopsis Barnacle goose Breeds in Arctic Russia, eastern Greenland and northern Europe; wintering in northern and northwestern Europe.
016 - CACKLING GOOSE (10-31-06) sloco, ca (8708309130).jpg Branta hutchinsii – formerly included in B. canadensis Cackling goose North America, northern Canada and Alaska

Two species have been described from subfossil remains found in the Hawaiian Islands, where they became extinct in prehistoric times:

Similar but hitherto undescribed remains are known from on Kauaʻi and Oʻahu.

The relationships of the enigmatic Geochen rhuax, formerly known only from parts of a single bird's skeleton damaged due to apparently dying in a lava flow, were long unresolved. After reexamination of the subfossil material and comparisons with other subfossil bones from the island of Hawaiʻi assigned to the genus Branta, it was redescribed as Branta rhuax in 2013. [6] While a presumed relation between B. rhuax and the shelducks, proposed by Lester Short in 1970, [7] has thus been refuted, bones of a shelduck-like bird have been found more recently on Kaua‘i.[ citation needed ] Whether this latter anatid was indeed a shelduck is presently undetermined.

Similarly, two bones found on Oʻahu indicate the erstwhile presence of a gigantic waterfowl on this island. Its relationships relative to this genus and the moa-nalos, enormous goose-like dabbling ducks, are completely undeterminable at present.

Early fossil record

Several fossil species of Branta have been described. Since the true geese are hardly distinguishable by anatomical features, the allocation of these to this genus is somewhat uncertain. A number of supposed prehistoric grey geese have been described from North America, partially from the same sites as species assigned to Branta. Whether these are correctly assigned – meaning that the genus Anser was once much more widespread than today and that it coexisted with Branta in freshwater habitat which it today does only most rarely – is not clear. Especially in the case of B. dickeyi and B. howardae, doubts have been expressed about its correct generic assignment.[ citation needed ]

The former "Branta" minuscula is now placed with the prehistoric American shelducks, Anabernicula . [10] On the other hand, a goose fossil from the Early-Middle Pleistocene of El Salvador is highly similar to Anser and given its age and biogeography it is likely to belong to that genus or Branta. [14]

Footnotes

  1. The nēnē, which is aberrant in many respects, has no white on the head or neck and fairly little black, being quite similar to the swan goose in the color pattern of these areas, and was thus formerly assigned to the monotypic genus Nesochen. The swan goose, a grey goose, also has a black bill, but its reddish-orange legs indicate its actual relationships.

Related Research Articles

<span class="mw-page-title-main">Goose</span> Common name for a group of waterfowl

A goose is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera Anser and Branta. Some members of the Tadorninae subfamily are commonly called geese, but are not considered "True Geese" taxonomically. More distantly related members of the family Anatidae are swans, most of which are larger than true geese, and ducks, which are smaller.

<span class="mw-page-title-main">Swan</span> Tribe of large water birds

Swans are birds of the genus Cygnus within the family Anatidae. The swans' closest relatives include the geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe Cygnini. Sometimes, they are considered a distinct subfamily, Cygninae.

<span class="mw-page-title-main">Loon</span> Family of birds

Loons or divers are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus Gavia, family Gaviidae and order Gaviiformes.

<span class="mw-page-title-main">Nene (bird)</span> Species of bird

The nene, also known as the nēnē or the Hawaiian goose, is a species of bird endemic to the Hawaiian Islands. The nene is exclusively found in the wild on the islands of Oahu, Maui, Kauaʻi, Molokai, and Hawaiʻi. In 1957, it was designated as the official state bird of the state of Hawaiʻi.

<span class="mw-page-title-main">Greylag goose</span> Species of bird

The greylag goose or graylag goose is a species of large goose in the waterfowl family Anatidae and the type species of the genus Anser. It has mottled and barred grey and white plumage and an orange beak and pink legs. A large bird, it measures between 74 and 91 centimetres in length, with an average weight of 3.3 kilograms. Its distribution is widespread, with birds from the north of its range in Europe and Asia often migrating southwards to spend the winter in warmer places, although many population are resident, even in the north. It is the ancestor of most breeds of domestic goose, having been domesticated at least as early as 1360 BCE. The genus name and specific epithet are from anser, the Latin for "goose".

<span class="mw-page-title-main">Red-breasted goose</span> Species of bird

The red-breasted goose is a brightly marked species of goose in the genus Branta from Eurasia. It is currently classified as vulnerable by the IUCN.

<span class="mw-page-title-main">Anatidae</span> Biological family of water birds

The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating on the water surface, and in some cases diving in at least shallow water. The family contains around 174 species in 43 genera.

<span class="mw-page-title-main">Anseriformes</span> Order of water birds

Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae, Anseranatidae, and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves. Due to their aquatic nature, most species are web-footed though this one is not.

<span class="mw-page-title-main">Anserinae</span> Subfamily of birds

The Anserinae are a subfamily in the waterfowl family Anatidae. It includes the swans and the true geese. Under alternative systematical concepts, it is split into two subfamilies, the Anserinae contain the geese and the ducks, while the Cygninae contain the swans.

<i>Tadorna</i> Genus of birds

The shelducks, most species of which are found in the genus Tadorna, are a group of large birds in the Tadorninae subfamily of the Anatidae, the biological family that includes the ducks and most duck-like waterfowl such as the geese and swans.

<span class="mw-page-title-main">Common shelduck</span> Species of bird

The common shelduck is a waterfowl species of the shelduck genus, Tadorna. It is widespread and common in the Euro-Siberian region of the Palearctic, mainly breeding in temperate and wintering in subtropical regions; in winter, it can also be found in the Maghreb.

<i>Anas</i> Genus of birds

Anas is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was split into four separate genera. The genus now contains 31 living species. The name Anas is the Latin for "duck".

<span class="mw-page-title-main">Ross's goose</span> Species of bird

Ross's goose, formerly Chen rossii, is a white goose with black wingtips and a relatively short neck. It is the smallest of the three white geese that breed in North America. It is similar in appearance to a white-phase snow goose, but about 40% smaller. Other differences from the snow goose are that the bill is smaller in proportion to its body and lacks "black lips". Like snow geese, Ross's geese may exhibit a darker "blue" phase or morph, though this is extremely rare.

<span class="mw-page-title-main">Blue-winged goose</span> Species of bird

The blue-winged goose is a waterfowl species which is endemic to Ethiopia. It is the only member of the genus Cyanochen.

<i>Anser</i> (bird) Genus of birds

Anser is a waterfowl genus that includes the grey geese and the white geese. It belongs to the true goose and swan subfamily of Anserinae under the family of Anatidae. The genus has a Holarctic distribution, with at least one species breeding in any open, wet habitats in the subarctic and cool temperate regions of the Northern Hemisphere in summer. Some also breed farther south, reaching into warm temperate regions. They mostly migrate south in winter, typically to regions in the temperate zone between the January 0 °C (32 °F) and 5 °C (41 °F) isotherms.

<i>Chendytes</i> Extinct species of bird

Chendytes lawi or Laws Duck is an extinct, goose-sized flightless marine duck, once common on the California coast, the California Channel Islands, and possibly southern Oregon. It lived in the Pleistocene and survived into the Holocene. It appears to have gone extinct at about 450–250 BCE. The youngest direct radiocarbon date from a Chendytes bone fragment dates to 770–400 BCE and was found in an archeological site in Ventura County. Its remains have been found in fossil deposits and in early coastal archeological sites. Archeological data from coastal California show a record of human exploitation of Chendytes lawi for at least 8,000 years. It was probably driven to extinction by hunting, animal predation, and loss of habitat. Chendytes bones have been identified in archaeological assemblages from 14 coastal sites, including two on San Miguel Island and 12 in mainland localities. Hundreds of Chendytes bones and egg shells found in Pleistocene deposits on San Miguel Island have been interpreted as evidence that some of these island fossil localities were nesting colonies, one of which Guthrie dated to about 12,000 14C years. There is nothing in the North American archaeological record indicating a span of exploitation for any megafaunal genus remotely as long as that of Chendytes.

<i>Branta rhuax</i> Extinct species of goose

Branta rhuax, the giant Hawaiʻi goose, is an extinct goose endemic to the island of Hawaiʻi. It was initially described as the monotypic genus Geochen, but then reassigned to Branta by Storrs L. Olson in 2013 after reexamination of the subfossil material.

<i>Coragyps occidentalis</i> Extinct species of New World vulture

Coragyps occidentalis, the Pleistocene black vulture, is an extinct species of New World vulture that lived throughout North and South America during the Pleistocene. It was formerly thought to be the ancestor to the modern black vulture, but is now thought to have evolved from it; the modern black vulture is paraphyletic with respect to it.

References

  1. Scopoli, Giovanni Antonio (1769). Annus Historico-Naturalis (in Latin). Vol. Part 1. Lipsiae [Leipzig]: Sumtib. C.G. Hilscheri. p. 67.
  2. Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p.  77. ISBN   978-1-4081-2501-4.
  3. Mayr, Ernst; Cottrell, G. William, eds. (1979). Check-List of Birds of the World. Vol. 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 440.
  4. Ottenburghs, J.; Megens, H.-J.; Kraus, R.H.S.; Madsen, O.; van Hooft, P.; van Wieren, S.E.; Crooijmans, R.P.M.A.; Ydenberg, R.C.; Groenen, M.A.M.; Prins, H.H.T. (2016). "A tree of geese: A phylogenomic perspective on the evolutionary history of True Geese". Molecular Phylogenetics and Evolution. 101: 303–313. doi:10.1016/j.ympev.2016.05.021. PMID   27233434.
  5. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2020). "Screamers, ducks, geese & swans". IOC World Bird List Version 10.2. International Ornithologists' Union. Retrieved 7 October 2020.
  6. Olson, Storrs L. (2013). "Hawaii's first fossil bird: history, geological age, and taxonomic status of the extinct goose Geochen rhuax Wetmore (Aves: Anatidae)". Proceedings of the Biological Society of Washington. Washington: Biological Society of Washington. 126 (2): 161–168. doi:10.2988/0006-324x-126.2.161. S2CID   86807390 . Retrieved 13 July 2013.
  7. 1 2 3 4 Short, Lester L. (1970). "A new anseriform genus and species from the Nebraska Pliocene" (PDF). Auk . 87 (3): 537–543. doi:10.2307/4083796. JSTOR   4083796.
  8. Miller, Loye (1924). "Branta dickeyi from the McKittrick Pleistocene" (PDF). The Condor. 26 (5): 178–180. doi:10.2307/1363171. JSTOR   1363171.
  9. 1 2 3 Miller, Loye (1944). "Some Pliocene Birds from Oregon and Idaho" (PDF). The Condor. 46 (1): 25–32. doi:10.2307/1364248. JSTOR   1364248.
  10. 1 2 3 4 5 6 Short, Lester L. (1969). "A new genus and species of gooselike swan from the Pliocene of Nebraska". American Museum Novitates (2369). hdl:2246/2579.
  11. 1 2 Miller, Alden H.; Ashley, James F. (1934). "Goose Footprints on a Pliocene Mud-flat" (PDF). The Condor. 36 (4): 178–179. doi:10.2307/1363416. JSTOR   1363416.
  12. Howard, Hildegarde (1931). "Pliocene Bird Remains from Santa Barbara, California" (PDF). The Condor. 33 (1): 30–31. doi:10.2307/1363932. JSTOR   1363932.
  13. Miller, Loye (1931). "Bird Remains from the Kern River Pliocene of California" (PDF). The Condor. 33 (2): 70–72. doi:10.2307/1363312. JSTOR   1363312.
  14. Cisneros, Juan Carlos (2005). "New Pleistocene vertebrate fauna from El Salvador" (PDF). Revista Brasileira de Paleontologia (in English and Portuguese). 8 (3): 239–255. doi: 10.4072/rbp.2005.3.09 .

Further reading