Anser | |
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Pair of greylag geese, Anser anser | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Anseriformes |
Family: | Anatidae |
Tribe: | Anserini Vigors 1825 |
Genus: | Anser Brisson, 1760 |
Type species | |
Anas anser [1] | |
Species | |
and see text | |
Synonyms | |
Chen Boie, 1822 (but see text) Contents |
Anser is a waterfowl genus that includes the grey geese and the white geese. It belongs to the true goose and swan subfamily of Anserinae under the family of Anatidae. [2] The genus has a Holarctic distribution, with at least one species breeding in any open, wet habitats in the subarctic and cool temperate regions of the Northern Hemisphere in summer. Some also breed farther south, reaching into warm temperate regions. They mostly migrate south in winter, typically to regions in the temperate zone between the January 0 °C (32 °F) and 5 °C (41 °F) isotherms.
The genus contains 11 living species. [2]
The species of this genus span nearly the whole range of true goose shapes and sizes. The largest are the bean, greylag and swan geese at up to around 4 kg (9 lb) in weight (with domestic forms far exceeding this), and the smallest are the lesser white-fronted and Ross's geese, which ranges from about 1.3 to 2.3 kg (3–5 lb). [3]
All have legs and feet that are pink, or orange, and bills that are pink, orange, or black. All have white under- and upper-tail coverts, and several have some extent of white on their heads. The neck, body and wings are grey or white, with black or blackish primary—and also often secondary—remiges (pinions). The three species of "white geese" (emperor, snow and Ross's geese) were formerly treated as a separate genus Chen, but are now generally included in Anser. The closely related "black" geese in the genus Branta differ in having black legs, and generally darker body plumage. [3]
The genus Anser was introduced by the French zoologist Mathurin Jacques Brisson in 1760. [4] The name comes from the Latin word anser meaning "goose" [5] used as the specific epithet for the greylag goose (Anas anser) introduced by Linnaeus in 1758, that epithet was repeated to become its generic name as the type species. [6] [7]
The evolutionary relationships between Anser geese have been difficult to resolve because of their rapid radiation during the Pleistocene and frequent hybridization. [8] [9] In 2016 Ottenburghs and colleagues published a study that established the phylogenetic relationships between the species by comparing exonic DNA sequences. [10]
Anser |
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The genus contains 11 species: [2]
Image | Scientific name | Common name | Distribution |
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Anser indicus | Bar-headed goose | Breeds in highlands of Central Asia; winters in South Asia, Myanmar and southern China; introduced in Europe | |
Anser canagicus | Emperor goose | Near the Pacific coast in Alaska, Russian Far East and Canada | |
Anser rossii | Ross's goose | Breeds in northern Canada and Alaska; winters in contiguous United States and northern Mexico | |
Anser caerulescens | Snow goose | Breeds in northern Canada, Alaska and Greenland; winters in contiguous United States and northern Mexico | |
Anser anser | Greylag goose | Europe, Asia and North Africa | |
Anser cygnoides | Swan goose | Breeds in Mongolia, northernmost China and southeastern Russia; winters in southeastern China | |
Anser fabalis | Taiga bean goose | Breeds in Russia, Finland, Norway and Sweden; winters in Europe, and Central and East Asia | |
Anser serrirostris | Tundra bean goose | Breeds in northern Russia; winters in Europe, and Central and East Asia | |
Anser brachyrhynchus | Pink-footed goose | Breeds in Iceland, Svalbard and Greenland; winters in northwestern Europe | |
Anser albifrons | Greater white-fronted goose | Breeds in northern Canada, Alaska, Greenland and northern Russia; winters in contiguous United States, northern Mexico, Europe, East Asia, Iraq and near the Caspian Sea | |
Anser erythropus | Lesser white-fronted goose | Breeds in northern Russia, Finland, Norway and Sweden; winters in East Asia, near the Caspian Sea, and in southeastern and northwestern Europe | |
The following white geese were separated as the genus Chen. Most ornithological works now include Chen within Anser, [11] [12] [13] [14]
Some authorities also treat some subspecies as distinct species (notably the tundra bean goose [15] [16] ) or as likely future species splits (notably the Greenland white-fronted goose). [17]
Numerous fossil species have been allocated to this genus. As the true geese are near-impossible to assign osteologically to genus, this must be viewed with caution. It can be assumed with limited certainty that European fossils from known inland sites belong into Anser. As species related to the Canada goose have been described from the Late Miocene onwards in North America too, sometimes from the same localities as the presumed grey geese, it casts serious doubt on the correct generic assignment of the supposed North American fossil geese. [18] [19] [20] Heterochen = Anser pratensis seems to differ profoundly from other species of Anser and might be placed into a different genus; alternatively, it might have been a unique example of a grey goose adapted for perching in trees. [a] [b]
The Maltese swan Cygnus equitum was occasionally placed into Anser, and Anser condoni is a synonym of Cygnus paloregonus . [18] A goose fossil from the Early-Middle Pleistocene of El Salvador is highly similar to Anser. [21] Given its age it is likely to belong to an extant genus, and biogeography indicates Branta as other likely candidate.
?Anser scaldii Beneden 1872 nomen nudum (Late Miocene of Antwerp, Belgium) may be a shelduck.
Two species in the genus are of major commercial importance, having been domesticated as poultry: European domesticated geese are derived from the greylag goose, and Chinese and some African domesticated geese are derived from the swan goose.
Most species are hunted to a greater or lesser extent; in some areas, some populations are threatened by over-hunting and habitat loss. Although most species are not considered threatened by the IUCN, the lesser white-fronted goose and swan goose are listed as Vulnerable and the emperor goose is near-threatened. [22] [23] [24]
Other species have benefited from reductions in hunting since the late 19th and early 20th centuries, with most species in western Europe and North America showing marked increases in response to protection[ citation needed ]. In some cases, this has led to conflicts with farming, when large flocks of geese graze crops in the winter.[ citation needed ]
A goose is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera Anser and Branta. Some members of the Tadorninae subfamily are commonly called geese, but are not considered "true geese" taxonomically. More distantly related members of the family Anatidae are swans, most of which are larger than true geese, and ducks, which are smaller.
Swans are birds of the genus Cygnus within the family Anatidae. The swans' closest relatives include the geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe Cygnini. Sometimes, they are considered a distinct subfamily, Cygninae.
The greylag goose or graylag goose is a species of large goose in the waterfowl family Anatidae and the type species of the genus Anser. It has mottled and barred grey and white plumage and an orange beak and pink legs. A large bird, it measures between 74 and 91 centimetres in length, with an average weight of 3.3 kilograms. Its distribution is widespread, with birds from the north of its range in Europe and Asia often migrating southwards to spend the winter in warmer places, although many populations are resident, even in the north. It is the ancestor of most breeds of domestic goose, having been domesticated at least as early as 1360 BCE. The genus name and specific epithet are from anser, the Latin for "goose".
The greater white-fronted goose is a species of goose, closely related to the smaller lesser white-fronted goose. The greater white-fronted goose is migratory, breeding in northern Canada, Alaska, Greenland and Russia, and winters farther south in North America, Europe and Asia. It is named for the patch of white feathers bordering the base of its bill: albifrons comes from the Latin albus "white" and frons "forehead". In the United Kingdom and Ireland, it has been known as the white-fronted goose; in North America it is known as the greater white-fronted goose, and this name is also increasingly adopted internationally. Even more distinctive are the salt-and-pepper markings on the breast of adult birds, which is why the goose is colloquially called the "specklebelly" in North America.
The lesser white-fronted goose is a goose closely related to the larger greater white-fronted goose. It breeds in the northernmost Palearctic, but it is a scarce breeder in Europe, with a reintroduction attempt in Fennoscandia.
The snow goose is a species of goose native to North America. Both white and dark morphs exist, the latter often known as blue goose. Its name derives from the typically white plumage. The species was previously placed in the genus Chen, but is now typically included in the "gray goose" genus Anser.
The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating on the water surface, and, in some cases, diving in at least shallow water. The family contains around 174 species in 43 genera.
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae, Anseranatidae, and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves, the clade consisting of all other modern birds except the galliformes and paleognaths. Due to their aquatic nature, most species are web-footed.
The Anserinae are a subfamily in the waterfowl family Anatidae. It includes the swans and the true geese. Under alternative systematical concepts, it is split into two subfamilies, the Anserinae contain the geese and the ducks, while the Cygninae contain the swans.
The Oxyurini are a tribe of the duck subfamily of birds, the Anatinae. It has been subject of considerable debate about its validity and circumscription. Some taxonomic authorities place the group in its own subfamily, the Oxyurinae. Most of its members have long, stiff tail feathers which are erected when the bird is at rest, and relatively large, swollen bills. Though their relationships are still enigmatic, they appear to be closer to swans and true geese than to the typical ducks. The highest diversity is found in the warmer parts of the Americas, but at least one species occurs in a major part of the world.
The swan goose is a large goose with a natural breeding range in inland Mongolia, Northeast China, and the Russian Far East. It is migratory and winters mainly in central and eastern China. Vagrant birds are encountered in Japan and Korea, and more rarely in Kazakhstan, Laos, coastal Siberia, Taiwan, Thailand and Uzbekistan.
The magpie goose is the sole living representative species of the family Anseranatidae. This common waterbird is found in northern Australia and southern New Guinea. As the species is prone to wandering, especially when not breeding, it is sometimes recorded outside its core range. The species was once also widespread in southern Australia but disappeared from there largely due to the drainage of the wetlands where the birds once bred. Due to their importance to Aboriginal people as a seasonal food source, as subjects of recreational hunting, and as a tourist attraction, their expansive and stable presence in northern Australia has been "ensured [by] protective management".
Anseranatidae, the magpie-geese, is a biological family of waterbirds. The only living species, the magpie goose, is a resident breeder in northern Australia and in southern New Guinea.
The black geese of the genus Branta are waterfowl belonging to the true geese and swans subfamily Anserinae. They occur in the northern coastal regions of the Palearctic and all over North America, migrating to more southerly coasts in winter, and as resident birds in the Hawaiian Islands. Alone in the Southern Hemisphere, a self-sustaining feral population derived from introduced Canada geese is also found in New Zealand.
Anas is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was split into four separate genera. The genus now contains 31 living species. The name Anas is the Latin for "duck".
Ross's goose, formerly Chen rossii, is a white goose with black wingtips and a relatively short neck. It is the smallest of the three white geese that breed in North America. It is similar in appearance to a white-phase snow goose, but about 40% smaller. Other differences from the snow goose are that the bill is smaller in proportion to its body and lacks "black lips". Like snow geese, Ross's geese may exhibit a darker "blue" phase or morph, though this is extremely rare.
Pygmy geese are a group of very small "perching ducks" in the genus Nettapus which breed in the Old World tropics. They are the smallest of all wildfowl. As the "perching ducks" are a paraphyletic group, they need to be placed elsewhere. The initially assumed relationship with the dabbling duck subfamily Anatinae has been questioned, and it appears they form a lineage in an ancient Gondwanan radiation of waterfowl, within which they are of unclear affinities. An undescribed fossil species from the late Hemphillian of Jalisco, central Mexico, has also been identified from the distal end of a tarsometatarsus. It is only record of the genus in the New World.
A bird hybrid is a bird that has two different species as parents. The resulting bird can present with any combination of characteristics from the parent species, from totally identical to completely different. Usually, the bird hybrid shows intermediate characteristics between the two species. A "successful" hybrid is one demonstrated to produce fertile offspring. According to the most recent estimates, about 16% of all wild bird species have been known to hybridize with one another; this number increases to 22% when captive hybrids are taken into account. Several bird species hybridize with multiple other species. For example, the mallard is known to interbreed with at least 40 different species. The ecological and evolutionary consequences of multispecies hybridization remain to be determined.
The tundra bean goose is a goose that breeds in northern Siberia. This and the taiga bean goose are recognised as separate species by the American Ornithological Society and International Ornithologists' Union, but are considered a single species by other authorities. It is migratory and winters further south in Asia. The taiga and tundra bean goose diverged about 2.5 million years ago and established secondary contact ca. 60,000 years ago, resulting in extensive gene flow.