Orthothecida

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Orthothecida
Temporal range: Early Cambrian – Early Devonian [1]
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Clade: Brachiozoa
Class: Hyolitha
Order: Orthothecida
Marek, 1966
Families

The orthothecids are one of the two hyolith orders. [2]

Marek diagnoses the order thus: Conchs with a flat or concave ventral surface — opercula with large, flat cardinal processes but without clavicles – tightly sigmoidal, sediment-filled intestine – helens absent.

Sometimes the Circothecidae and Tetrathecidae are split out into a separate order 'Circothecida', which is defined by the bottom surface not being flat, the cardinal processes being pronounced, and a circular rim sometimes showing hints of differentiation into clavicles. [3]

Internal taxonomy

Marek [4] gives the following diagnoses:

Kouchinsky [5] lists the following taxonomic criteria:

Elsewhere [6] :243 is stated:

Additional genera are not assigned to a family:

Related Research Articles

Hyolitha Palaeozoic lophophorates with small conical shells

Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic Era. They are lophophorates, a group which includes the brachiopods.

Clavicle Long bone that serves as a strut between the scapula and the sternum

The clavicle, or collarbone, is a slender, S-shaped flat bone approximately 6 inches (15 cm) long that serves as a strut between the shoulder blade and the sternum (breastbone). There are two clavicles, one on the left and one on the right. The clavicle is the only long bone in the body that lies horizontally. Together with the shoulder blade, it makes up the shoulder girdle. It is a touchable bone, and in people who have less fat in this region, the location of the bone is clearly visible, as it creates a bulge in the skin. It receives its name from the Latin clavicula, because the bone rotates along its axis like a key when the shoulder is abducted. The clavicle is the most commonly fractured bone. It can easily be fractured by impacts to the shoulder from the force of falling on outstretched arms or by a direct hit.

Acromion Bony process on the scapula (shoulder blade)

In human anatomy, the acromion is a bony process on the scapula. Together with the coracoid process it extends laterally over the shoulder joint. The acromion is a continuation of the scapular spine, and hooks over anteriorly. It articulates with the clavicle to form the acromioclavicular joint.

Tibia Long bone of the lower leg

The tibia, also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates, and it connects the knee with the ankle bones. The tibia is found on the medial side of the leg next to the fibula and closer to the median plane or centre-line. The tibia is connected to the fibula by the interosseous membrane of leg, forming a type of fibrous joint called a syndesmosis with very little movement. The tibia is named for the flute tibia. It is the second largest bone in the human body next to the femur. The leg bones are the strongest long bones as they support the rest of the body.

Halkieriid Family of incertae sedis

The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.

Evolution of molluscs The origin and diversification of molluscs through geologic time

The evolution of the molluscs is the way in which the Mollusca, one of the largest groups of invertebrate animals, evolved. This phylum includes gastropods, bivalves, scaphopods, cephalopods, and several other groups. The fossil record of mollusks is relatively complete, and they are well represented in most fossil-bearing marine strata. Very early organisms which have dubiously been compared to molluscs include Kimberella and Odontogriphus.

Plectronocerida Extinct order of molluscs

Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.

Turtle shell

The turtle shell is a shield for the ventral and dorsal parts of turtles, completely enclosing all the vital organs of the turtle and in some cases even the head. It is constructed of modified bony elements such as the ribs, parts of the pelvis and other bones found in most reptiles. The bone of the shell consists of both skeletal and dermal bone, showing that the complete enclosure of the shell probably evolved by including dermal armor into the rib cage.

Helcionellid Extinct order of molluscs

Helcionellid or Helcionelliformes is an order of small fossil shells that are universally interpreted as molluscs, though no sources spell out why this taxonomic interpretation is preferred. These animals are first found about 540 to 530 million years ago in the late Nemakit-Daldynian age, which is the earliest part of the Cambrian period. A single species persisted to the Early Ordovician. These fossils are component of the small shelly fossils (SSF) assemblages.

<i>Scenella tenuistriata</i>

Scenella tenuistriata is an extinct species generally classified as a mollusc or hydrozoan in the family Scenellidae.

Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.

<i>Kurtziella rhysa</i> Species of gastropod

Kurtziella rhysa is a species of sea snail, a marine gastropod mollusk in the family Mangeliidae.

<i>Turbo setosus</i> Species of gastropod

Turbo setosus, common name the rough turban, is a species of sea snail, marine gastropod mollusk in the family Turbinidae.

Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.

<i>Fordilla</i> Extinct genus of bivalves

Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.

The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.

Stenothecoida is a taxon of bivalved fossils from the Early to middle Cambrian period. They look a bit like brachiopods or bivalve molluscs.

Mobergella is a millimetric Lower Cambrian shelly fossil of unknown affinity, usually preserved in phosphate and particularly well known from Swedish strata, where it is diagnostic of lowermost Cambrian rocks. Originally interpreted as a monoplacophoran, the circular, cap-shaped shell resembles a hyolith operculum, with concentric rings on its upper surface, and seven pairs of internal muscle scars. It is never found in association with a conch, and its affinity therefore remains undetermined. Nevertheless, its heavy musculature does seem to indicate that it functioned as an operculum.

Fordilloidea Extinct superfamily of bivalves

Fordilloidea is an extinct superfamily of early bivalves containing two described families, Fordillidae and Camyidae and the only superfamily in the order Fordillida. The superfamily is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. Fordillidae currently contains two genera, Fordilla and Pojetaia each with up to three described species while Camyidae only contains a single genus Camya with one described species, Camya asy. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.

<i>Theta spicea</i> Species of gastropod

Theta spicea is a species of sea snail, a marine gastropod mollusk in the family Raphitomidae.

References

  1. Malinky, J. M. (2009). "Permian Hyolithida from Australia: The Last of the Hyoliths?". Journal of Paleontology. 83: 147–152. doi:10.1666/08-094R.1. S2CID   85924056.
  2. Malinky, J.M.; Skovsted, C.B (2004). "Hyoliths and small shelly fossils from the Lower Cambrian of North−East Greenland". Acta Palaeontologica Polonica. 49 (4): 551–578.
  3. http://palaeontologia.pan.pl/Archive/1994-53-1-2_213-252_52-56.pdf [ bare URL PDF ]
  4. "MACHAERIDIANS, CHITONS, and CONCHIFERAN: MOLLUSCS of the MOJCZA LIMESTONE" (PDF).{{cite journal}}: Cite journal requires |journal= (help)
  5. Kouchinsky, A. V. (2000). "Skeletal microstructures of hyoliths from the Early Cambrian of Siberia". Alcheringa. 24 (2): 65–81. doi:10.1080/03115510008619525. S2CID   140660142.
  6. Lipps, Jere H; Signor, Philip W (2013-11-21). Origin and Early Evolution of the Metazoa. ISBN   9781489924278.
  7. Valent, M. (2012). "Two new orthothecids from the Cambrian of the Barrandian area (Hyolitha, Skryje-Týřovice Basin, Czech Republic)". Bulletin of Geosciences. 87 (2): 241–248. doi: 10.3140/bull.geosci.1142 .
  8. Sun, Haijing; Smith, Martin R.; Zeng, Han; Zhao, Fangchen; Li, Guoxiang; Zhu, Maoyan (2018). "Hyoliths with pedicles illuminate the origin of the brachiopod body plan". Proceedings of the Royal Society B: Biological Sciences. 285 (1887): 20181780. doi:10.1098/rspb.2018.1780. PMC   6170810 . PMID   30257914.