Orthothecida

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Orthothecida
Temporal range: Early Cambrian – Early Devonian [1]
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Brachiopoda
Class: Hyolitha
Order: Orthothecida
Marek, 1966
Families

The orthothecids are one of the two hyolith orders. [2]

Marek diagnoses the order thus: Conchs with a flat or concave ventral surface — opercula with large, flat cardinal processes but without clavicles – tightly sigmoidal, sediment-filled intestine – helens absent.

Sometimes the Circothecidae and Tetrathecidae are split out into a separate order 'Circothecida', which is defined by the bottom surface not being flat, the cardinal processes being pronounced, and a circular rim sometimes showing hints of differentiation into clavicles. [3]

Internal taxonomy

Marek [3] gives the following diagnoses:

Kouchinsky [4] lists the following taxonomic criteria:

Elsewhere [5] :243 is stated:

Additional genera are not assigned to a family:

Related Research Articles

<span class="mw-page-title-main">Hyolitha</span> Palaeozoic lophophorates with small conical shells

Hyoliths are animals with small conical shells, known from fossils from the Palaeozoic era. They are at least considered as being lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs.

<span class="mw-page-title-main">Clavicle</span> Long bone that serves as a strut between the scapula and the sternum

The clavicle, collarbone, or keybone is a slender, S-shaped long bone approximately 6 inches (15 cm) long that serves as a strut between the shoulder blade and the sternum (breastbone). There are two clavicles, one on the left and one on the right. The clavicle is the only long bone in the body that lies horizontally. Together with the shoulder blade, it makes up the shoulder girdle. It is a palpable bone and, in people who have less fat in this region, the location of the bone is clearly visible. It receives its name from Latin clavicula 'little key' because the bone rotates along its axis like a key when the shoulder is abducted. The clavicle is the most commonly fractured bone. It can easily be fractured by impacts to the shoulder from the force of falling on outstretched arms or by a direct hit.

<span class="mw-page-title-main">Ulna</span> Medial bone from forearm

The ulna or ulnar bone is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the radius, the forearm's other long bone. Longer and thinner than the radius, the ulna is considered to be the smaller long bone of the lower arm. The corresponding bone in the lower leg is the fibula.

<span class="mw-page-title-main">Acromion</span> Bony process on the scapula (shoulder blade)

In human anatomy, the acromion is a bony process on the scapula. Together with the coracoid process, it extends laterally over the shoulder joint. The acromion is a continuation of the scapular spine, and hooks over anteriorly. It articulates with the clavicle to form the acromioclavicular joint.

<span class="mw-page-title-main">Meniscus (liquid)</span> Curve in a liquids surface due to adhesion to the container walls

In physics, the meniscus is the curve in the upper surface of a liquid close to the surface of the container or another object, produced by surface tension.

<span class="mw-page-title-main">Tibia</span> Leg bone in vertebrates

The tibia, also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates ; it connects the knee with the ankle. The tibia is found on the medial side of the leg next to the fibula and closer to the median plane. The tibia is connected to the fibula by the interosseous membrane of leg, forming a type of fibrous joint called a syndesmosis with very little movement. The tibia is named for the flute tibia. It is the second largest bone in the human body, after the femur. The leg bones are the strongest long bones as they support the rest of the body.

<span class="mw-page-title-main">Molding (decorative)</span> Class of decorative elements in the ornamentation

Moulding, or molding, also coving, is a strip of material with various profiles used to cover transitions between surfaces or for decoration. It is traditionally made from solid milled wood or plaster, but may be of plastic or reformed wood. In classical architecture and sculpture, the moulding is often carved in marble or other stones. In historic architecture, and some expensive modern buildings, it may be formed in place with plaster.

<span class="mw-page-title-main">Halkieriid</span> Family of extinct molluscs

The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.

<i>Haplophrentis</i> Extinct genus of Cambrian organisms

Haplophrentis is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called helens protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts.

<i>Haliotis corrugata</i> Species of gastropod

The pink abalone, scientific name Haliotis corrugata, is a species of large edible sea snail, a marine gastropod mollusk in the family Haliotidae, the abalones.

<span class="mw-page-title-main">Helcionellid</span> Extinct order of molluscs

Helcionellid or Helcionelliformes is an order of small fossil shells that are universally interpreted as molluscs, though no sources spell out why this taxonomic interpretation is preferred. These animals are first found about 540 to 530 million years ago in the late Nemakit-Daldynian age, which is the earliest part of the Cambrian period. A single species persisted to the Early Ordovician. These fossils are component of the small shelly fossils (SSF) assemblages.

Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.

<i>Turbo cornutus</i> Species of gastropod

Turbo cornutus, common name the horned turban, is a species of sea snail, marine gastropod mollusk in the family Turbinidae.

Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.

<i>Fordilla</i> Extinct genus of bivalves

Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.

The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.

Mobergella is a millimetric Lower Cambrian shelly fossil of unknown affinity, usually preserved in phosphate and particularly well known from Swedish strata, where it is diagnostic of lowermost Cambrian rocks. Originally interpreted as a monoplacophoran, the circular, cap-shaped shell resembles a hyolith operculum, with concentric rings on its upper surface, and seven pairs of internal muscle scars. It is never found in association with a conch, and its affinity therefore remains undetermined. Nevertheless, its heavy musculature does seem to indicate that it functioned as an operculum.

<span class="mw-page-title-main">Hyolithida</span> Extinct order of molluscs

The Hyolithida are lophophorates, one of the two orders of hyolithid, the other being the Orthothecida. Most of our knowledge of the hyolithids comes from studies on the Hyolithida. Both orders had an operculum that was not hinged to the conch. However, the Hyolithida are distinct from the Orthothecida in having additional paired, curved, whiskerlike appendages. The Hyolithida were probably bottom feeders living in shallow water, and had tentacules.

Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.

<span class="mw-page-title-main">Strophomenata</span> Extinct class of marine lamp shells

Strophomenata is an extinct class of brachiopods in the subphylum Rhynchonelliformea.

References

  1. Malinky, J. M. (2009). "Permian Hyolithida from Australia: The Last of the Hyoliths?". Journal of Paleontology. 83: 147–152. doi:10.1666/08-094R.1. S2CID   85924056.
  2. Malinky, J.M.; Skovsted, C.B (2004). "Hyoliths and small shelly fossils from the Lower Cambrian of North−East Greenland". Acta Palaeontologica Polonica. 49 (4): 551–578.
  3. 1 2 "Machaeridians, Chitons, and Conchiferan Molluscs of the Mojcza limestone" (PDF). Archived from the original (PDF) on 2017-10-12.
  4. Kouchinsky, A. V. (2000). "Skeletal microstructures of hyoliths from the Early Cambrian of Siberia". Alcheringa. 24 (2): 65–81. doi:10.1080/03115510008619525. S2CID   140660142.
  5. Lipps, Jere H; Signor, Philip W (2013-11-21). Origin and Early Evolution of the Metazoa. ISBN   9781489924278.
  6. Valent, M. (2012). "Two new orthothecids from the Cambrian of the Barrandian area (Hyolitha, Skryje-Týřovice Basin, Czech Republic)". Bulletin of Geosciences. 87 (2): 241–248. doi: 10.3140/bull.geosci.1142 .
  7. Sun, Haijing; Smith, Martin R.; Zeng, Han; Zhao, Fangchen; Li, Guoxiang; Zhu, Maoyan (2018). "Hyoliths with pedicles illuminate the origin of the brachiopod body plan". Proceedings of the Royal Society B: Biological Sciences. 285 (1887): 20181780. doi:10.1098/rspb.2018.1780. PMC   6170810 . PMID   30257914.