The history of Crete goes back to the 7th millennium BC, preceding the ancient Minoan civilization by more than four millennia. The palace-based Minoan civilization was the first civilization in Europe.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Pliocene, and expanded into Eurasia during the Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus, the latter of which has been suggested to be the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition. Some modern populations of Asian elephants have also undergone size reduction on islands to a lesser degree, resulting in populations of pygmy elephants.
Palaeoloxodon falconeri is an extinct species of dwarf elephant from the Middle Pleistocene of Sicily and Malta. It is amongst the smallest of all dwarf elephants at only 1 metre (3.3 ft) in height. A member of the genus Palaeoloxodon, it derived from a population of the mainland European straight-tusked elephant.
The Cypriot mouse is a species of mouse endemic to Cyprus. Its primary habitat seems to be the vineyards and fields of the Troödos Mountains region.
Candiacervus is an extinct genus of deer native to Pleistocene Crete. Due to a lack of other herbivores, the genus underwent an adaptive radiation, filling niches occupied by other taxa on the mainland. Due to the small size of Crete, some species underwent insular dwarfism, the smallest species, C. ropalophorus, stood about 40 centimetres (16 in) at the shoulders when fully grown, while other species were relatively large and comparable in size to mainland deer species. Some species are noted for their peculiar, elongate club-shaped antlers, though other species have more normal antlers.
The Cyprus dwarf hippopotamus or Cypriot pygmy hippopotamus is an extinct species of hippopotamus that inhabited the island of Cyprus from the Pleistocene until the early Holocene. The 200-kilogram (440 lb) Cyprus dwarf hippo was roughly the same size as the extant pygmy hippopotamus. Unlike the modern pygmy hippo, the Cyprus dwarf became small through the process of insular dwarfism. H. minor is the smallest hippopotamus of all known insular hippopotamuses. It is estimated to have measured 76 cm (2.5 ft) tall and 121 cm (4.0 ft) long.
Hippopotamus melitensis is an extinct hippopotamus from Malta. It lived during Middle-Late Pleistocene. It probably descended from Hippopotamus pentlandi from Sicily, which in turn probably descended from the common hippopotamus. Like Hippopotamus pentlandi, Hippopotamus melitensis is substantially smaller in size than H. amphibius as a result of insular dwarfism, having an estimated mass of approximately 900 kg, which is smaller than the 1100 kg estimated for H. pentlandi. The diet of H. melitensis is suggested to have been more generalist than Hippopotamus amphibius, likely as a result of limited resource diversity and lack of competition, as the only other large herbivore on the island was the dwarf elephant Palaeoloxodon mnaidriensis. The majority of findings of this species are from Għar Dalam, a cave on Malta famous for its Pleistocene fossil deposits.
Hippopotamus pentlandi is an extinct species of hippopotamus from Sicily, known from the late Middle Pleistocene to early Late Pleistocene. It is the largest of the insular dwarf hippos known from the Pleistocene of the Mediterranean, "at most 20% smaller than the mainland forms", with an estimated body mass of approximately 1100 kg. It is suggested that it arrived in Sicily between 250,000 and 150,000 years ago, probably descending from the modern hippopotamus, with an origin from Hippopotamus antiquus being less likely. In comparison to those species, the muzzle was shorter, the occipital and nasal regions were more developed, the mastoid process was enlarged, and the dental row was shortened, and the condyle of the mandible is low. In comparison to H. amphibius, the orbits are also elevated. It was present in Sicily until at least the latest Middle Pleistocene around 120 kya, and was probably extinct by the beginning of Marine Isotope Stage 4. Contemporaneous species include the dwarf elephant Palaeoloxodon mnaidriensis, the aurochs, red deer, steppe bison, fallow deer, wild boar, brown bear, wolves, red foxes, cave hyena and cave lions. Its diet was likely grazing dominated, similar to that of modern H. amphibius. It is probably ancestral to Hippopotamus melitensis from Malta, which is substantially smaller than H. pentlandi.
Palaeoloxodon cypriotes is an extinct species of dwarf elephant that inhabited the island of Cyprus during the Late Pleistocene. Remains comprise 44 molars, found in the north of the island, seven molars discovered in the south-east, a single measurable femur and a single tusk among very sparse additional bone and tusk fragments. The molars support derivation from the large straight-tusked elephant (Palaeoloxodon antiquus). The species is presumably derived from the older, larger P. xylophagou from the late Middle Pleistocene which reached the island presumably during a Pleistocene glacial maximum when low sea levels allowed a low probability sea crossing between Cyprus and Asia Minor. During subsequent periods of isolation the population adapted within the evolutionary mechanisms of insular dwarfism, which the available sequence of molar fossils confirms to a certain extent. The fully developed Palaeoloxodon cypriotes weighed not more than 200 kg (440 lb) and had a height of around 1 m (3.28 ft). The species became extinct around 12,000 years ago, around the time humans first colonised Cyprus.
Hippopotamus creutzburgi, the Cretan dwarf hippopotamus, is an extinct species of hippopotamus from the island of Crete. It lived on the island from the Early Pleistocene to early Middle Pleistocene, and probably descended from Hippopotamus antiquus. It was considerably smaller than H. antiquus, weighing approximately 400 kilograms (880 lb). It was one of only two large herbivores on the island during its existence, alongside the dwarf mammoth Mammuthus creticus, with large predators being absent, the only other species of mammal present on the island being the giant rat Kritimys. It is known from abundant remains collected from the Katharo basin in the eastern uplands of Crete, approximately 1,100–1,200 metres (3,600–3,900 ft) above sea level, as well as much rarer remains found in coastal caves. Analysis of its limbs suggests that it was more adapted to terrestrial locomotion than living hippopotamus, primarily walking on its hooves rather than its footpads as in living hippopotamus, and capable of traversing the rugged terrain of Crete. Analysis of its teeth suggests that it had a grazing diet, similar to modern Hippopotamus amphibius. The previous suggestion that the species can be divided into two subspecies is not supported by modern research.
Palaeoloxodon mnaidriensis is an extinct species of dwarf elephant belonging to the genus Palaeoloxodon, native to the Siculo-Maltese archipelago during the late Middle Pleistocene and Late Pleistocene. It is derived from the European mainland straight-tusked elephant.
The wildlife of Cyprus includes its flora and fauna and their natural habitats. Cyprus has a rich flora and a diverse fauna albeit with relatively few mammals. Like most modern countries, the natural habitats in Cyprus have been steadily disappearing, currently retaining only 20% of its original habitat due to rapid urbanization, usage of forests for commercial purposes, tourism and various other reasons. One of the features of Cyprus' habitats is the wild and sharp differences in elevations and habitats on the island as well as climate, all of which supply a diverse habitat for an array of fauna and flora. Terra Cypria was established as a trust in 1992 to conserve the Cypriot environment and its biodiversity.
Mammuthus creticus, or the Cretan dwarf mammoth, is an extinct species of dwarf mammoth endemic to Crete. It is only known from a single locality near Chania, which probably dates to the Early or Middle Pleistocene. With a shoulder height of about 1 m and a weight of about 180 kg, it was the smallest mammoth that ever existed. Its size reduction was the result of insular dwarfism, where the body size of large mammals on islands reduces as the result of decreased food availability, predation and competition. It was originally described as Elephas creticus by Dorothea Bate in 1907. After DNA research published in 2006, it was proposed to rename Elephas creticus into Mammuthus creticus. Others proposed to rename all the described specimens of larger size under the new subspecies name Elephas antiquus creutzburgi. A 2007 study criticised the results of the 2006 DNA study, showing that the DNA research was likely flawed. However, morphological data supports a placement in Mammuthus. It probably derived from Mammuthus meridionalis, or less likely, Mammuthus rumanus. It was one of only three mammal species native to Crete during the Early Pleistocene and the early Middle Pleistocene, alongside the dwarf hippopotamus Hippopotamus creutzburgi and the giant rat Kritimys.
Leithia is an extinct genus of giant dormice from the Pleistocene of the Mediterranean islands of Malta and Sicily. It is considered an example of island gigantism. Leithia melitensis is the largest known species of dormouse, living or extinct, being twice the size of any other known species.
Elephas beyeri is an extinct species of dwarf elephant belonging to the Elephantidae family of the Middle Pleistocene. It was named after the anthropologist H. Otley Beyer. The type specimen was discovered on Cabarruyan Island in The Philippines but has since been lost.
The Crete Mediterranean forests is a terrestrial ecoregion that encompasses the Greek island of Crete.
Kritimys, also known as the Cretan giant rat is an extinct genus of murid rodent that was endemic to the island of Crete during the Early and Middle Pleistocene. There are two known species, K. kiridus from the Early-Mid Pleistocene, and its descendant K. catreus from the Middle Pleistocene. It is suggested to be closely related to and probably derived from Praomys. As with most island rodents, Kritimys was larger than its mainland relatives, with its size increasing over time, with K. catreus esimated to weigh 518 grams (1.142 lb), around 6.7 times the weight of its mainland ancestor, an example of island gigantism. The temporal range of the genus is considered to define the regional Kritimys biozone, during which time there were only two other species of mammal native to the island, a species of dwarf mammoth, Mammuthus creticus and the dwarf hippopotamus Hippopotamus creutzburgi. It became extinct during the late Middle Pleistocene, following the arrival of the Mus bateae-minotaurus lineage to the island, exhibiting a decrease in size shortly before its extinction.
The Cretan otter(Lutrogale cretensis) is an extinct otter that was endemic to Crete during the Pleistocene.
Mus minotaurus is an extinct species of mouse native to Crete during the Late Pleistocene-Holocene. It descended from a Mus musculus -like ancestor that arrived on Crete during the late Middle Pleistocene, replacing Kritimys, a large rat-like rodent that inhabited Crete during the Early and Middle Pleistocene. Both Kritimys and the ancestor of Mus minotaurus, Mus bateae are found together in Stavrós Cave. The Mus batae-minotaurus lineage shows a tendency to increase in size with time, an example of island gigantism, with Mus minotaurus being one of the largest known members of the genus Mus, with a body mass of approximately 54 grams. It was likely heavily predated upon by the extinct endemic Cretan owl, as evidenced by the abundance of its remains found in owl pellets. It inhabited the island alongside a species of elephant, the radiation of endemic Candiacervus deer, the Cretan otter, and the Cretan shrew. Mus minotaurus became extinct sometime during the Holocene epoch, with its remains apparently being found in Neolithic and early Bronze Age sites on the island. Its extinction may have been due to competition with the closely related house mouse introduced to the island by humans during the early Bronze Age.
