A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly late Miocene.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition.
The pygmy mammoth or Channel Islands mammoth is an extinct species of dwarf mammoth native to the northern Channel Islands off the coast of southern California during the Late Pleistocene. It was descended from the Columbian mammoth of mainland North America, which are suggested to have colonised the islands around 250-150,000 years ago. At only 1.72–2.02 m (5.6–6.6 ft) tall at the shoulder, it was around 17% the size of its mainland ancestor. The species became extinct around 13,000 years ago, co-inciding with major environmental change and the arrival of humans in the Channel islands.
The Columbian mammoth is an extinct species of mammoth that inhabited North America from southern Canada to Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian steppe mammoths that colonised North America during the Early Pleistocene around 1.5–1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
Mammuthus meridionalis, sometimes called the southern mammoth, is an extinct species of mammoth native to Eurasia, including Europe, during the Early Pleistocene, living from around 2.5 million years ago to 800,000 years ago.
Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.
Mammuthus lamarmorai is a species of dwarf mammoth which lived during the late Middle and Late Pleistocene on the island of Sardinia in the Mediterranean. It has been estimated to have had a shoulder height of around 1.4 metres (4.6 ft). Remains have been found across the western part of the island.
The Cypriot pygmy hippopotamus is an extinct species of dwarf hippopotamus that inhabited the island of Cyprus from the Pleistocene until the early Holocene.
Mammuthus africanavus is a species of mammoth known from remains spanning the Late Pliocene-Early Pleistocene found in Central and North Africa in the countries of Chad, Morocco, Tunisia and Algeria. It was originally described by Camille Arambourg in 1952 based on remains found around Lake Ichkeul in north Tunisia as a species of Elephas. Some specimens from this sample may genuinely represent Elephas rather than Mammuthus, though the holotype has been argued to likely represent a true mammoth. Some authors have argued that the species should be placed in Loxodonta, reflecting the difficulty in distinguishing the teeth of early elephantids. It is distinguished from the earlier Mammuthus subplanifrons by having a higher number of ridges/lamellae on the teeth, which display a greater parallelity, the molars being more hypsodont, with the molars having a greater amount of cementum and thinner enamel, and the molar plates exhibit closer spacing.
Palaeoloxodon cypriotes is an extinct species of dwarf elephant that inhabited the island of Cyprus during the Late Pleistocene. The species became extinct around 12,000 years ago, around the time humans first colonised Cyprus.
Dorothea Minola Alice Bate, also known as Dorothy Bate, was a Welsh palaeontologist and pioneer of archaeozoology. Her life's work was to find fossils of recently extinct mammals with a view to understanding how and why giant and dwarf forms evolved.
Primelephas is a genus of Elephantinae that existed during the Miocene and Pliocene epochs. The name of the genus suggests 'first elephant'. These primitive elephantids are thought to be the common ancestor of Mammuthus, the mammoths, and the closely allied genera Elephas and Loxodonta, the Asian and African elephants, diverging some 4-6 million years ago. It had four tusks, which is a trait not shared with its descendants, but common in earlier proboscideans. The type species, Primelephas gomphotheroides, was described by Vincent Maglio in 1970, with the specific epithet indicating the fossil specimens were gomphothere-like. Primelephas korotorensis is the only other species to be assigned to the genus. All fossils found of the Primelephas have been found in Africa, primarily in modern day Chad, Tanzania, Kenya, Ethiopia, and Uganda.
Hippopotamus creutzburgi, the Cretan dwarf hippopotamus, is an extinct species of hippopotamus from the island of Crete. It lived on the island from the Early Pleistocene to early Middle Pleistocene, and probably descended from Hippopotamus antiquus. It was considerably smaller than H. antiquus, weighing approximately 400 kilograms (880 lb). It was one of only two large herbivores on the island during its existence, alongside the dwarf mammoth Mammuthus creticus, with large predators being absent, the only other species of mammal present on the island being the giant rat Kritimys. It is known from abundant remains collected from the Katharo basin in the eastern uplands of Crete, approximately 1,100–1,200 metres (3,600–3,900 ft) above sea level, as well as much rarer remains found in coastal caves. Analysis of its limbs suggests that it was more adapted to terrestrial locomotion than living hippopotamus, primarily walking on its hooves rather than its footpads as in living hippopotamus, and capable of traversing the rugged terrain of Crete. Analysis of its teeth suggests that it had a grazing diet, similar to modern Hippopotamus amphibius. The previous suggestion that the species can be divided into two subspecies is not supported by modern research.
Palaeoloxodon creutzburgi is an extinct species of elephant known from the Middle-Late Pleistocene of Crete. It is a descendant of the large mainland species Palaeoloxodon antiquus. It is known from localities across the island. P. chaniensis from Stylos and in Vamos cave, Chania, west Crete is considered to be a junior synonym of P. creutzburgi. It had undergone insular dwarfism, being approximately 40% of the size of its mainland ancestor, and was around the size of the living Asian elephant. It lived alongside the radiation of Candiacervus deer endemic to the island, the mouse Mus batae-minotaurus, the Cretan otter, and the Cretan shrew.
The Crete Mediterranean forests is a terrestrial ecoregion that encompasses the Greek island of Crete.
Kritimys, also known as the Cretan giant rat is an extinct genus of murid rodent that was endemic to the island of Crete during the Early and Middle Pleistocene. There are two known species, K. kiridus from the Early-Mid Pleistocene, and its descendant K. catreus from the Middle Pleistocene. It is suggested to be closely related to and probably derived from Praomys. As with most island rodents, Kritimys was larger than its mainland relatives, with its size increasing over time, with K. catreus estimated to weigh 518 grams (1.142 lb), around 6.7 times the weight of its mainland ancestor, an example of island gigantism. The temporal range of the genus is considered to define the regional Kritimys biozone, during which time there were only two other species of mammal native to the island, a species of dwarf mammoth, Mammuthus creticus and the dwarf hippopotamus Hippopotamus creutzburgi. It became extinct during the late Middle Pleistocene, following the arrival of the Mus bateae-minotaurus lineage to the island, exhibiting a decrease in size shortly before its extinction.
