Elephas Temporal range: Pliocene to present | |
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Asian elephant | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Proboscidea |
Family: | Elephantidae |
Tribe: | Elephantini |
Genus: | Elephas Linnaeus, 1758 |
Type species | |
Elephas maximus | |
Species | |
For others, see text | |
Synonyms | |
Hypselephas |
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. [1] Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly the late Miocene.
Species of Elephas have distinct bossing of the parieto-occipital region of the skull. The premaxillae bones containing the tusks are tapered. [2]
Relationships of living and extinct elephantids based on DNA, after Palkopoulou et al. 2018. [3]
Elephantidae |
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Asian elephants share a closer common ancestry with mammoths (genus Mammuthus) than they do with African elephants (Loxodonta). [4] The oldest species attributed to the genus Elephas is E. nawataensis from the Late Miocene-Early Pliocene of Kenya, though the validity of this species and its relationship to Elephas has been doubted. [5] The oldest species widely attributed to the genus, Elephas ekorensis is known from the early-mid Pliocene (5–4.2 million years ago) of East Africa, [6] though the attribution of this species to Elephas has been questioned, due to a lack of shared morphological features with later Elephas species. [7] The oldest record of the genus outside of Africa is Elephas planifrons which is known from the Late Pliocene of the Indian subcontinent, around 3.6 million years ago. [8] However, the placement of Elephas planifrons within the genus has also been questioned. [9] The earliest fossils of the ancestor of the modern Asian elephant, Elephas hysudricus date to the beginning of the Pleistocene, around 2.6 million years ago, with remains found on the Indian subcontinent. [10] Modern Asian elephants had evolved from E. hysrudicus by the Late Pleistocene. [11]
The scientific name Elephas was proposed by Carl Linnaeus in 1758 who described the genus and an elephant from Ceylon. [12] The genus is assigned to the proboscidean family Elephantidae and is made up of one living and seven extinct species: [13]
The following Asian elephants were proposed as extinct subspecies, but are now considered synonymous with the Indian elephant: [1]
The following Elephas species are extinct:
While formerly assigned to this genus, Elephas recki , Elephas namadicus , the straight-tusked elephant E. antiquus and the dwarf elephants E. falconeri and E. cypriotes are now placed in the separate genus Palaeoloxodon, which is more closely related to African elephants. [23] However, some material historically assigned to Elephas recki , such as Elephas recki atavus, may be closely related to true Elephas, rather than to Palaeoloxodon [24] "Elephas" celebensis is now placed in Stegoloxodon . [25]
A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene until about 4,000 years ago, with mammoth species at various times inhabiting Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
A mastodon is a member of the genus Mammut, which, strictly defined, was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 27–25 million years ago, during the Oligocene.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae.
Palaeoloxodon recki, often known by the synonym Elephas recki, is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. One of the largest known elephant species, mature fully grown bulls on average had a shoulder height of 4 metres (13 ft) and a weight of 13 tonnes (29,000 lb). Straight-tusked elephants likely lived very similarly to modern elephants, with herds of adult females and juveniles and solitary adult males. The species was primarily associated with temperate and Mediterranean woodland and forest habitats, flourishing during interglacial periods, when its range would extend across Europe as far north as Great Britain and eastwards into Russia. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestor of numerous species of dwarf elephants that inhabited islands in the Mediterranean.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Mammuthus meridionalis, sometimes called the southern mammoth, is an extinct species of mammoth native to Eurasia, including Europe, during the Early Pleistocene, living from around 2.5 million years ago to 800,000 years ago.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the early Middle to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
Mammuthus africanavus is a species of mammoth known from remains spanning the Late Pliocene-Early Pleistocene found in Central and North Africa in the countries of Chad, Morocco, Tunisia and Algeria. It was originally described by Camille Arambourg in 1952 based on remains found around Lake Ichkeul in north Tunisia as a species of Elephas. Some specimens from this sample may genuinely represent Elephas rather than Mammuthus, though the holotype has been argued to likely represent a true mammoth. Some authors have argued that the species should be placed in Loxodonta, reflecting the difficulty in distinguishing the teeth of early elephantids. It is distinguished from the earlier Mammuthus subplanifrons by having a higher number of ridges/lamellae on the teeth, which display a greater parallelity, the molars being more hypsodont, with the molars having a greater amount of cementum and thinner enamel, and the molar plates exhibit closer spacing.
Palaeoloxodon naumanni is an extinct species of elephant belonging to the genus Palaeoloxodon that was native to the Japanese archipelago during the Middle to Late Pleistocene around 330,000 to 24,000 years ago. It is named after the German geologist Heinrich Edmund Naumann who first described remains of the species in the 19th century, with the species sometimes being called Naumann's elephant. Fossils attributed to P. naumanni are also known from China, though the status of these specimens is unresolved, and some authors regard them as belonging to separate species.
Elephas ekorensis is an extinct species of elephant. Fossils have been found in East Africa. They date as far back as the Early Pliocene age, between 5 and 4.2 million years ago. It is the earliest species placed in the genus Elephas. It has been suggested to have been a grazer or mixed feeder. Its placement in the genus of Elephas has been questioned, as the teeth are similar to those of the contemporaneous Loxodonta adaurora. A number of specimens assigned to it likely actually belong to other species.
Palaeoloxodonjolensis is an extinct species of elephant. The type specimen is located in the National Museum of Natural History in Paris. It is either considered the descendant species or last evolutionary stage of Palaeoloxodon recki in Africa. It is only known from isolated molars. The species is known from remains found across Africa, which are largely poorly dated to approximately the late Middle Pleistocene to Late Pleistocene, with some authors suggesting an exclusively late Middle Pleistocene age, as the only well dated specimens of the species are over 130,000 years old. Like P. recki, they are thought to have been dedicated grazers, being distinguished from earlier P. recki by having increased hypsodonty and enamel folding, with the plates being thicker along the long axis of the tooth. Following the extinction of the species, it was largely replaced by the modern African bush elephant.
Elephas planifrons is an extinct species of elephant, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent.
Stegoloxodon is an extinct genus of dwarf elephant known from the Early Pleistocene of Indonesia. It contains two species, S. indonesicus from Java, and S. celebensis from Sulawesi. Its relationship with other elephants is uncertain.
Phanagoroloxodon is a genus of extinct elephant. It is known from one species, Phanagoroloxodon mammontoides, which is described from a partial skull from Russia, of probable Late Pliocene-Early Pleistocene age.