Javan elephant | |
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Partial fossil tooth of an Asian elephant collected in Cipeundeuy, West Bandung Regency, West Java, possibly of a Javan elephant | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Proboscidea |
Family: | Elephantidae |
Genus: | Elephas |
Species: | |
Subspecies: | E. m. sondaicus |
Trinomial name | |
Elephas maximus sondaicus Deraniyagala, 1953 [1] |
The Javan elephant (Elephas maximus sondaicus) was proposed by Paules Edward Pieris Deraniyagala in 1953, based on an illustration of a carving on the Buddhist monument of Borobudur in Java. He thought that the Asian elephant (Elephas maximus) had indeed existed on the island and had gone extinct. [2] It may be considered synonymous with the Sumatran elephant (E. maximus sumatranus). [1]
Fossils of the Asian elephant have been found in Pleistocene deposits on Java. The question of when elephants became extinct on Java is unsettled. Chinese chronicles contemporary with the period of Hindu influence in Java recorded that Javan kings rode on elephants, and that Java exported ivory to China. As elephants were, at least occasionally, transported by ship, it is possible that the elephants in Java during the period of Hindu influence had been imported from India. [3]
A tradition in the northeastern part of Borneo holds that the Borneo elephants that currently live in the wild there, and the elephants that formerly lived in the wild on the neighboring island of Sulu, are descended from elephants from Java that were presented by the "Raja of Java" (perhaps, the leader of Majapahit) to Rajah Baguinda of Sulu at the end of the 14th century. Another tradition holds that elephants were presented to the Sultan of Sulu by the East India Company in 1750. The lack of a fossil record of elephants on Borneo has been cited as support for the theory that elephants have been recently introduced to Borneo.
Fernando, et al., found that the elephants on Borneo have been genetically isolated from other Asian elephant populations for approximately 300,000 years, and, assuming that any elephants introduced into Borneo by the East India Company were from India, Sumatra, or peninsular Malaya, concluded that the elephants in Borneo are indigenous, representing a colonization during the Pleistocene. The Earl of Cranbrook, et al. conclude that either an indigenous occupation of Borneo since the Pleistocene, or a recent introduction from Java, are plausible for the origin for the Borneo elephants. If the Borneo elephants are descended from Javan elephants, then the Javan elephants presumably would have also been genetically distinct from other Asian elephant populations. [4] [5]
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
The Javan rhinoceros, Javan rhino, Sunda rhinoceros or lesser one-horned rhinoceros is a critically endangered member of the genus Rhinoceros, of the rhinoceros family, Rhinocerotidae, and one of the five remaining extant rhinoceros species across South Asia and Africa. The Javan rhinoceros is one of the smallest rhinoceros species, along with the Sumatran, or "hairy", rhinoceros. They are superficially similar to Indian rhinos, as they have plate-like, "armored" protective skin folds, but are slightly smaller, at just 3.1–3.2 m (10–10 ft) long and 1.4–1.7 m (4.6–5.6 ft) tall, on average. The heaviest specimens weigh around 2,300 kg/2.3 tonnes, similar to a black rhinoceros. However, unlike the long and potentially lethal horns of the black or white rhinoceroses of Africa, the Javan species' single, somewhat blunted horn is usually shorter than 25 cm (9.8 in).
The Asian elephant, also known as the Asiatic elephant, is a species of elephant distributed throughout the Indian subcontinent and Southeast Asia, from India in the west to Borneo in the east, and Nepal in the north to Sumatra in the south. Three subspecies are recognised—E. m. maximus, E. m. indicus and E. m. sumatranus. The Asian elephant is characterised by its long trunk with a single finger-like processing; large tusks in males; laterally folded large ears but smaller in contrast to African elephants; and wrinkled grey skin. The skin is smoother than African elephants and may be depigmented on the trunk, ears or neck. Adult males average 4 tonnes in weight, and females 2.7 t.
The Sumatran rhinoceros, also known as the Sumatran rhino, hairy rhinoceros or Asian two-horned rhinoceros, is a rare member of the family Rhinocerotidae and one of five extant species of rhinoceros; it is the only extant species of the genus Dicerorhinus. It is the smallest rhinoceros, although it is still a large mammal; it stands 112–145 cm (44–57 in) high at the shoulder, with a head-and-body length of 2.36–3.18 m and a tail of 35–70 cm (14–28 in). The weight is reported to range from 500–1,000 kg (1,100–2,200 lb), averaging 700–800 kg (1,540–1,760 lb). Like both African species, it has two horns; the larger is the nasal horn, typically 15–25 cm (5.9–9.8 in), while the other horn is typically a stub. A coat of reddish-brown hair covers most of the Sumatran rhino's body.
Sundaland is a biogeographical region of Southeast Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It includes Bali, Borneo, Java, and Sumatra in Indonesia, and their surrounding small islands, as well as the Malay Peninsula on the Asian mainland.
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly the late Miocene.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition.
Ratnapura is a major city in Sri Lanka. It is the capital city of Sabaragamuwa Province, as well as the Ratnapura District, and is a traditional centre for the Sri Lankan gem trade. It is located on the Kalu Ganga in south-central Sri Lanka, some 101 km (63 mi) southeast of the country's capital, Colombo. Ratnapura is also spelled as Rathnapura.
The Sumatran elephant is one of three recognized subspecies of the Asian elephant, and native to the Indonesian island of Sumatra. In 2011, IUCN upgraded the conservation status of the Sumatran elephant from endangered to critically endangered in its Red List as the population had declined by at least 80% during the past three generations, estimated to be about 75 years. The subspecies is preeminently threatened by habitat loss, degradation and fragmentation, and poaching; over 69% of potential elephant habitat has been lost within the last 25 years. Much of the remaining forest cover is in blocks smaller than 250 km2 (97 sq mi), which are too small to contain viable elephant populations.
The Sri Lankan elephant is native to Sri Lanka and one of three recognised subspecies of the Asian elephant. It is the type subspecies of the Asian elephant and was first described by Carl Linnaeus under the binomial Elephas maximus in 1758. The Sri Lankan elephant population is now largely restricted to the dry zone in the north, east and southeast of Sri Lanka. Elephants are present in Udawalawe National Park, Yala National Park, Lunugamvehera National Park, Wilpattu National Park and Minneriya National Park but also live outside protected areas. It is estimated that Sri Lanka has the highest density of elephants in Asia. Human-elephant conflict is increasing due to conversion of elephant habitat to settlements and permanent cultivation.
The Indian elephant is one of three extant recognized subspecies of the Asian elephant, native to mainland Asia. The species is smaller than the African elephant species with a convex back and the highest body point on its head. The species exhibits significant sexual dimorphism with a male reaching an average shoulder height of about 3.2 m (10 ft) and weighing up to 5,400 kg (11,900 lb) whereas a female reaches an average shoulder height of about 2.54 m (8.3 ft) and weighs up to 4,160 kg (9,170 lb). It has a broader skull with a concave forehead, two large laterally folded ears and a large trunk. It has grey colored smooth skin with four large legs and a long tail.
The Borneo elephant, also called the Bornean elephant or the Borneo pygmy elephant, is a subspecies of Asian elephant (Elephas maximus) that inhabits northeastern Borneo, in Indonesia and Malaysia. Its origin remains the subject of debate. A definitive subspecific classification as Elephas maximus borneensis awaits a detailed range-wide morphometric and genetic study. In 2024, the Borneo elephant has been listed as Endangered on the IUCN Red List as the population has declined by at least 50% over the last three generations, estimated to be 60–75 years. It is pre-eminently threatened by loss, degradation and fragmentation of habitat.
The Syrian or Western Asiatic elephant was the westernmost population of the Asian elephant, which went extinct in ancient times, with early human civilizations in the area utilizing the animals for their ivory, and possibly for warfare. Skeletal remains of E. m. asurus have been recorded in the Middle East, notably from Iran, Iraq, Syria, and Turkey, from periods dating between at least 1800 BC and likely 700 BC. Due to the lack of any Late Pleistocene or early to mid-Holocene records for Asian elephants in the region, there are suggestions that the elephants were anthropogenically introduced, or transported, there during the Bronze Age, though this is disputed.
Elephas hysudricus is an extinct elephant species known from the Pleistocene of Asia. It is thought to be ancestral to the living Asian elephant, from which it is distinguished by the molar teeth having a lower crown height and a lower lamellae number. Remains of the species are primarily known from the Indian subcontinent, with the most important remains coming from the Siwalik Hills. The oldest remains of the species in the Siwaliks are placed at around 2.6 million year ago at the beginning of the Early Pleistocene, with the youngest dates in the Siwaliks during the Middle Pleistocene around 0.6 million years ago, though it likely persisted on the subcontinent later than this based on remains found elsewhere.
Paulus Edward Pieris Deraniyagala (1900–1976) was a Sri Lankan paleontologist, zoologist, and artist.
Elephas hysudrindicus, commonly known also as the Blora elephant in Indonesia, is a species of extinct elephant from the Pleistocene of Java. It is anatomically distinct from the Asian elephant, the last remaining species of elephant under the genus Elephas. The species existed from around the end of the Early Pleistocene until the end of the Middle Pleistocene, when it was replaced by the modern Asian elephant in Java. It coexisted with the fellow proboscidean Stegodon trigonocephalus, as well as archaic humans belonging to the species Homo erectus.
The island of Borneo is located on the Sunda Shelf, which is an extensive region in Southeast Asia of immense importance in terms of biodiversity, biogeography and phylogeography of fauna and flora that had attracted Alfred Russel Wallace and other biologists from all over the world.
The Bornean tiger or Borneo tiger is possibly an extinct tiger population that lived on the island of Borneo in prehistoric times.