Elephas hysudrindicus Temporal range: Pleistocene | |
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Elephas hysudrindicus in the Bandung Geological Museum | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Proboscidea |
Family: | Elephantidae |
Genus: | Elephas |
Species: | †E. hysudrindicus |
Binomial name | |
†Elephas hysudrindicus Dubois, 1908 | |
Sunggun Archaeological Site in Blora, Java |
Elephas hysudrindicus, commonly known also as the Blora elephant in Indonesia (lit. Gajah Blora in Indonesian), is a species of extinct elephant from the Pleistocene of Java. It is anatomically distinct from the Asian elephant, the last remaining species of elephant under the genus Elephas . [1] [2] The species existed from around the end of the Early Pleistocene until the end of the Middle Pleistocene, when it was replaced by the modern Asian elephant in Java. It coexisted with the fellow proboscidean Stegodon trigonocephalus, as well as archaic humans belonging to the species Homo erectus . [3] [4]
When Eugène Dubois described the species in 1908, he failed to designate a holotype specimen. In 2017, the partial skull RGM.DUB 4968–4969 in the collections of the Naturalis Biodiversity Center, Leiden was designated the lectotype. It was excavated in Tinggang, Bojonegoro, and was Dubois' original fossil material. [2] It is considered to be closely related to the extinct Elephas hysudricus of mainland Asia, [5] which it is possibly descended from. [6]
A fossil was excavated from Sunggun archaeological site, Medalem, Kradenan Subregency, Blora in March 2009. It was excavated 13 kilometres west of the original Tinggang site [7] and was found almost completely intact (estimating about 90%), about a few feet under the dirt in a former sand quarry in the village. [8] [9] Both the skull and mandible were found to possess complete grinding molars, indicating that the individual was an adult specimen by time of death. [7] It was then brought to the Bandung Geological Museum which was then put to display. [10]
Isotopic analysis suggests a primarily C4 grazing dominated diet, similar to that inferred for Stegodon trigonocephalus, suggesting that they occupied open habitats on the island. [3]
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Java Man is an early human fossil discovered in 1891 and 1892 on the island of Java (Indonesia). Estimated to be between 700,000 and 1,490,000 years old, it was, at the time of its discovery, the oldest hominid fossil ever found, and it remains the type specimen for Homo erectus.
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly the late Miocene.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Homo floresiensis(), also known as "Flores Man" or "Hobbit", is an extinct species of small archaic humans that inhabited the island of Flores, Indonesia, until the arrival of modern humans about 50,000 years ago.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition.
Liang Bua is a limestone cave on the island of Flores, Indonesia, slightly north of the town of Ruteng in Manggarai Regency, East Nusa Tenggara. The cave demonstrated archaeological and paleontological potential in the 1950s and 1960s as described by the Dutch missionary and archaeologist Theodor L. Verhoeven.
Palaeoloxodon recki, often known by the synonym Elephas recki, is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related and ancestral to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki, P. recki ileretensis and Palaeoloxodon are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the Middle Pleistocene to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
Stegodon aurorae is an extinct species of Stegodon known from Early Pleistocene of Japan and possibly Taiwan. It appears to be an insular dwarf species, exhibiting smaller body size than its mainland ancestors.
Panthera tigris soloensis, known as the Ngandong tiger, is an extinct subspecies of the modern tiger species. It inhabited the Sundaland region of Indonesia during the Pleistocene epoch.
Mata Menge is an early Middle Pleistocene paleoanthropological site located in the Ola Bula Formation in the So'a Basin on the island of Flores, Indonesia. Lithic artefacts and hominin remains have been discovered at the site. The level of sophistication of the Mata Menge lithic artefacts is described as being 'simple'.
The region of Southeast Asia is considered a possible place for the evidence of archaic human remains that could be found due to the pathway between Australia and mainland Southeast Asia, where the migration of multiple early humans has occurred out of Africa. One of many pieces of evidence is of the early human found in central Java of Indonesia in the late 19th century by Eugene Dubois, and later in 1937 at Sangiran site by G.H.R. van Koenigswald. These skull and fossil materials are Homo erectus, named Pithecanthropus erectus by Dubois and Meganthropus palaeojavanicus by van Koenigswald. They were dated to c. 1.88 and 1.66 Ma, as suggested by Swisher et al. by analysis of volcanic rocks.
Duboisia santeng or Dubois' antelope is an extinct antelope-like bovid that was endemic to Indonesia during the Pleistocene. It went extinct during the Ionian stage of the Pleistocene, about 750.000 years ago. Duboisia santeng was first described by the Dutch paleoanthropologist and geologist Eugène Dubois in 1891.
Mececyon trinilensis, the Trinil dog, is an extinct canid species that lived on the island of Java in Indonesia during the Pleistocene.
Varanus hooijeri is an extinct species of medium-sized monitor lizard found in Liang Bua on Flores and possibly also Sumba, dating to the Late Pleistocene and Holocene.
Stegoloxodon is an extinct genus of dwarf elephant known from the Early Pleistocene of Indonesia. It contains two species, S. indonesicus from Java, and S. celebensis from Sulawesi. Its relationship with other elephants is uncertain.