The Proboscidea are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From the mid-Miocene onwards, most proboscideans were very large. The largest land mammal of all time may have been a proboscidean; Palaeoloxodon namadicus was up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, almost double the weight of several sauropods including Diplodocus carnegii and Apatosaurus louisae. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
A mastodon is any proboscidean belonging to the extinct genus Mammut that inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the Pleistocene 10,000 to 11,000 years ago. Mastodons lived in herds and were predominantly forest-dwelling animals that lived on a mixed diet obtained by browsing and grazing, somewhat similar to their distant relatives, modern elephants, but probably with greater emphasis on browsing.
Deinotherium was a large prehistoric relative of modern-day elephants that appeared in the Middle Miocene and survived until the Early Pleistocene. During that time, it changed very little. In life, it probably resembled modern elephants, except it had downward-curving tusks attached to the lower jaw.
Moeritherium is an extinct genus of primitive proboscideans. These prehistoric mammals are related to the elephant and, more distantly, sea cows and hyraxes. They lived during the Eocene epoch.
Stegodon, meaning "roofed tooth" because of the distinctive ridges on the animal's molars, is a genus of the extinct subfamily Stegodontinae of the order Proboscidea. It was assigned to the family Elephantidae, but has also been placed in the Stegodontidae. Stegodonts were present from 11.6 million years ago (Mya) to the late Pleistocene, with unconfirmed records of localized survival until 4,100 years ago. Fossils are found in Asian and African strata dating from the late Miocene; during the Pleistocene, they lived across large parts of Asia and East and Central Africa, and in Wallacea as far east as Timor.
Gomphotheres are any members of the diverse, extinct taxonomic family Gomphotheriidae. Gomphotheres were elephant-like proboscideans, but not belonging to the family Elephantidae. They were widespread in North America during the Miocene and Pliocene epochs, 12–1.6 million years ago (Mya). Some lived in parts of Eurasia, Beringia, and South America following the Great American Interchange. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, after these forms became extinct, the surviving gomphotheres had short jaws with either vestigal or no lower tusks (brevirostrine), looking very similar to modern elephants, an example of parallel evolution. Beginning after 2 Mya, they were gradually replaced by mammoths and mastodons in most of North America, with the last two genera, Cuvieronius persisting in southern North America and Notiomastodon having a wide range over most of South America, until the end of the Pleistocene, around 11,000 years ago.
Arsinoitherium is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the late Eocene and the early Oligocene of northern Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, A. giganteum, lived in Ethiopia about 27 million years ago.
Deinotheriidae is a family of prehistoric elephant-like proboscideans that lived during the Cenozoic era, first appearing in Africa, then spreading across southern Asia (Indo-Pakistan) and Europe. During that time, they changed very little, apart from growing much larger in size; by the late Miocene, they had become the largest land animals of their time. Their most distinctive features were the downward-curving tusks on the lower jaw.
Prodeinotherium is an extinct representative of the family Deinotheriidae that lived in Africa, Europe, and Asia in the early and middle Miocene. Prodeinotherium, meaning "before terrible beast", was first named in 1930, but soon after, the only species in it, P. hungaricum, was reassigned to Deinotherium. During the 1970s, however, the two genera were once again separated, with Prodeinotherium diagnosed to include Deinotherium bavaricum, Deinotherium hobleyi, and Deinotherium pentapotamiae, which were separated based on geographic location. The three species are from Europe, Africa, and Asia, respectively. However, because of usage of few characters to separate them, only one species, P. bavaricum, or many more species, including P. cuvieri, P. orlovii, and P. sinense may be possible.
Paraceratherium is an extinct genus of hornless rhinoceros. It is one of the largest terrestrial mammals that has existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. It is classified as a member of the hyracodont subfamily Indricotheriinae. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species A. bugtiense was originally placed.
The Columbian mammoth is an extinct species of mammoth that inhabited North America as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. It was one of the last in a line of mammoth species, beginning with Mammuthus subplanifrons in the early Pliocene. DNA studies show that the Columbian mammoth was a hybrid species between woolly mammoths and another lineage descended from steppe mammoths; the hybridization happened more than 420,000 years ago. The pygmy mammoths of the Channel Islands of California evolved from Columbian mammoths. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
Gomphotherium is an extinct genus of proboscids from the Neogene and early Pleistocene of Eurasia, Africa, North America and possibly also Asia.
Daouitherium is an extinct genus of early proboscideans that lived during the early Eocene some 55 million years ago in North Africa.
Numidotherium is an extinct genus of early proboscideans, discovered in 1984, that lived during the middle Eocene of North Africa some 46 million years ago. It was about 90-100 cm tall at the shoulder and weighed about 250-300 kg.
Phosphatherium escuillei is a basal proboscidean that lived from the Late Paleocene to the early stages of the Ypresian age until the early Thanetian some 56 million years ago in North Africa. Research has suggested that Phosphatherium existed during the Eocene period.
Dromicosuchus is an extinct genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It was found in Upper Triassic rocks of North Carolina, United States, and is known from a nearly complete skull and partial skeleton. This specimen is unusual in that it was found beneath the skeleton of a larger rauisuchian and has apparent bite damage, suggesting that it was attacked by the larger carnivore before both died and were buried together.
Palaeomastodon an extinct genus of Proboscidea. Palaeomastodon fossils have been found in Africa, lived some 36-35 million years ago. They are believed to be the ancestors of elephants or mastodons. Palaeomastodon lived in marshy semi aquatic swamps during the middle late Eocene to the early Oligocene. It may have used its upper pair of tusks for scraping bark off trees. Palaeomastodon was a very early form of the elephantidae and thus had a very short trunk.
Tetralophodon is an extinct elephantoid genus belonging to the family Anancidae.
Eritherium is an extinct genus of early Proboscidea found in the Ouled Abdoun basin, Morocco. It lived about 60 million years ago. It was first named by Emmanuel Gheerbrant in 2009 and the type species is Eritherium azzouzorum. Eritherium is the oldest, smallest and most primitive known elephant relative.
Ocepeia is an extinct genus of afrotherian mammal that lived in present-day Morocco during the middle Paleocene epoch, approximately 60 million years ago. First named and described in 2001, the type species is O. daouiensis from the Selandian stage of Morocco's Ouled Abdoun Basin. A second, larger species, O. grandis, is known from the Thanetian, a slightly younger stage in the same area. In life, the two species are estimated to have weighed about 3.5 kg (7.7 lb) and 10 kg (22 lb), respectively, and are believed to have been specialized leaf-eaters. The fossil skulls of Ocepeia are the oldest known afrotherian skulls, and the best-known of any Paleocene mammal in Africa.
