Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three living species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. In some cases, all members of the family can be referred to as elephants. They are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Moeritherium is an extinct genus of basal proboscideans from the Eocene of North and West Africa. The first specimen was discovered in strata from the Fayum fossil deposits of Egypt. It was named in 1901 by Charles William Andrews, who suggested that it was an early proboscidean, perhaps ancestral to mastodons, although subsequent workers considered it everything from a relative of manatees to a close relative of both clades' common ancestor. Currently, Moeritherium is seen as a proboscidean that, while fairly basal, predates the split between elephantiforms and deinotheres. Seven species have been named, though only three, are currently considered valid.
Deinotheriidae is a family of prehistoric elephant-like proboscideans that lived during the Cenozoic era, first appearing in Africa during the Oligocene then spreading across Europe and the lower latitudes of Asia during the Miocene epoch. Their most distinctive features were their lack of upper tusks and downward-curving tusks on the lower jaw.
Phenacodus is an extinct genus of mammals from the late Paleocene through middle Eocene, about 55 million years ago. It is one of the earliest and most primitive of the ungulates, typifying the family Phenacodontidae and the order Perissodactyla.
Phiomia, is an extinct genus of basal elephantiform proboscidean that lived in what is now Northern Africa during the Late Eocene to Early Oligocene some 37–30 million years ago. The type specimen of Phiomia, part of the mandible, was described in 1902 by Charles William Andrews and Hugh John Llewellyn. Unsure of its identity, they assigned it, tentatively, to the obsolete order Creodonta. Subsequently, it was recognised as a proboscidean. Briefly it was treated as a junior synonym of Palaeomastodon, but the two are regarded as separate genera. Though five species have been assigned to Phiomia over the years, only two, P. serridens and P. major, are currently recognised.
Numidotheriidae is an extinct family of primitive proboscideans that lived from the late Paleocene to the early Oligocene periods of North Africa.
Numidotherium is an extinct genus of early proboscideans, discovered in 1984, that lived during the middle Eocene of North Africa some 46 million years ago. It was about 90–100 cm (35–39 in) tall at the shoulder and weighed about 250–300 kg (550–660 lb).
Phosphatherium escuillei Gheerbrant, Sudre et Cappetta 1996 is a basal proboscidean that lived in Africa during the early Eocene, by 56-55 Ma. It is one of the earliest known proboscidean, together with Eritherium azzouzorum from the Selandian. It was found in phosphorites beds from the base of the Ypresian stage of the Ouled Abdoun Basin, which is best known for its exceptionally rich marine vertebrate fauna.
Barytherium is a genus of an extinct genus of large primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa. The type species is Barytherium grave, found at the beginning of the 20th century in Fayum, Egypt. Since then, more complete specimens have been found at Dor el Talha, Libya. More fossils were also discovered in 2011 in the Aidum area in Dhofar by Oman's Ministry of Heritage and Culture, which was named Barytherium omansi.
UA 8699 is a fossil mammalian tooth from the Cretaceous of Madagascar. A broken lower molar about 3.5 mm (0.14 in) long, it is from the Maastrichtian of the Maevarano Formation in northwestern Madagascar. Details of its crown morphology indicate that it is a boreosphenidan, a member of the group that includes living marsupials and placental mammals. David W. Krause, who first described the tooth in 2001, interpreted it as a marsupial on the basis of five shared characters, but in 2003 Averianov and others noted that all those are shared by zhelestid placentals and favored a close relationship between UA 8699 and the Spanish zhelestid Lainodon. Krause used the tooth as evidence that marsupials were present on the southern continents (Gondwana) as early as the late Cretaceous and Averianov and colleagues proposed that the tooth represented another example of faunal exchange between Africa and Europe at the time.
Eritherium is an extinct genus of early Proboscidea found in the Ouled Abdoun basin, Morocco. It lived about 60 million years ago. It was first named by Emmanuel Gheerbrant in 2009 and the type species is Eritherium azzouzorum. Eritherium is the oldest, smallest and most primitive known elephant relative.
Coloniatherium is a meridiolestid mammal from the Late Cretaceous of Argentina. The single species, Coloniatherium cilinskii, was a large member of the family Mesungulatidae.
Gandheralophus is an extinct genus of isectolophid perissodactyl mammal known from the Eocene of Pakistan, with two known species: G. minor and G. robustus, both described in 2012.
Megalochoerus is an extinct genus of large and long-legged pig-like animals from the Miocene of Africa.
Ocepeia is an extinct genus of afrotherian mammal that lived in present-day Morocco during the middle Paleocene epoch, approximately 60 million years ago. First named and described in 2001, the type species is O. daouiensis from the Selandian stage of Morocco's Ouled Abdoun Basin. A second, larger species, O. grandis, is known from the Thanetian, a slightly younger stage in the same area. In life, the two species are estimated to have weighed about 3.5 kg (7.7 lb) and 10 kg (22 lb), respectively, and are believed to have been specialized leaf-eaters. The fossil skulls of Ocepeia are the oldest known afrotherian skulls, and the best-known of any Paleocene mammal in Africa.
The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.
Arcanotherium is an extinct genus of early proboscidean belonging to the family Numidotheriidae that lived in North Africa during the late Eocene/early Oligocene interval.
Saloumia is an extinct genus of the order Proboscidea. It is one of the oldest members of the order and lived in the middle Eocene of Senegal. It is known only from a single molar, whose pronounced bumpy chewing surface indicates it is probably closely related to Moeritherium.
Dagbatitherium is an extinct genus of proboscideans. So far a single molar from the phosphate basins of Togo in West Africa has been found. The fossil dates to the Middle Eocene, around 47 million years ago. A striking feature of the tooth are the three pairs of cusps oriented transversely to the longitudinal axis of the tooth. This feature is found in more derived proboscideans, which are grouped in the Elephantiformes. For its age, Dagbatitherium is the earliest member of Elephantiformes to date. Furthermore, it is characterized by a low tooth crown and a humped occlusal pattern. The genus was described in 2021.
