Rhynchotherium

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Rhynchotherium
Temporal range: Miocene–Pliocene
Rhynchotherium.jpg
Restoration
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: Gomphotheriidae
Genus: Rhynchotherium
Falconer, 1868
Type species
Rhynchotherium falconeri
Osborn, 1923
Species
  • R. browni
  • R. edensis
  • R. falconeri
  • R. paredensis
  • R. simpsoni

Rhynchotherium is an extinct genus of proboscidea endemic to North America and Central America during the Miocene through Pliocene from 13.650 to 3.6 Ma, living for approximately 10 million years. [1]

Contents

This gomphothere had two tusks and may have evolved from Gomphotherium . [2]

Taxonomy

Rhynchotherium was first described in 1868 on the basis of a lower jaw from the Miocene of Tlaxcala, Mexico. [3] Later, the type species epithet R. tlascalae was erected for the jaw by Henry Fairfield Osborn in 1918. In 1921, a gomphothere skull from the Mt. Eden area of southern California was described as a subspecies of Trilophodon shepardi (a now-defunct combination for Mastodon shepardi), T. s. edensis, [4] but was subsequently reassigned to Rhynchotherium. [5] Other species subsequently assigned to Rhynchotherium included R. falconeri, [6] R. paredensis, R. browni, [7] and R. simpsoni. [8] It was the closest relative to Cuvieronius , and may be ancestral to it. [9]

Lucas and Morgan (2008) reviewed the taxonomy of Rhynchotherium and concluded that only R. edensis, R. falconeri, R. paredensis, R. browni, and R. simpsoni could be confidently referred to Rhynchotherium. [10] Because the genotype of Rhynchotherium is referable to Gomphotherium, the ICZN was petitioned to conserve the genus by designating R. falconeri as the type species, [11] which it did. [12]

Life restoration of Rhynchotherium falconeri, a North American species of the genus. Rhynchotherium falconeri.png
Life restoration of Rhynchotherium falconeri, a North American species of the genus.

Misassigned species

Phylogenetic position according to Mothé et al. (2016) [9]

Gomphotheriidae (Gomphotheres)

Gomphotherium

Gnathabelodon

Eubelodon

Brevirostrine clade

Stegomastodon

Sinomastodon

Notiomastodon

Rhynchotherium

Cuvieronius

Related Research Articles

<span class="mw-page-title-main">Proboscidea</span> Order of mammals including elephants

Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.

<span class="mw-page-title-main">Mastodon</span> Extinct genus of proboscideans

A mastodon is a member of the genus Mammut, which was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 27–25 million years ago, during the Oligocene.

<span class="mw-page-title-main">Mammutidae</span> Extinct family of mammals

Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae.

<i>Prodeinotherium</i> Extinct genus of proboscideans

Prodeinotherium is an extinct representative of the family Deinotheriidae that lived in Africa, Europe, and Asia in the early and middle Miocene. Prodeinotherium, meaning "before terrible beast", was first named in 1930, but soon after, the only species in it, P. hungaricum, was reassigned to Deinotherium. During the 1970s, however, the two genera were once again separated, with Prodeinotherium diagnosed to include Deinotherium bavaricum, Deinotherium hobleyi, and Deinotherium pentapotamiae, which were separated based on geographic location. The three species are from Europe, Africa, and Asia, respectively. However, because of usage of few characters to separate them, only one species, P. bavaricum, or many more species, including P. cuvieri, P. orlovii, and P. sinense may be possible.

<i>Gomphotherium</i> Extinct genus of elephant-like mammals

Gomphotherium is an extinct genus of gomphothere proboscidean from the Neogene of Eurasia, Africa and North America. It is the most diverse genus of gompothere, with over a dozen valid species. The genus is probably paraphyletic.

<i>Anancus</i> Genus of proboscideans

Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.

<i>Cuvieronius</i> Extinct genus of mammals

Cuvieronius is an extinct New World genus of gomphothere which ranged from southern North America to western South America during the Pleistocene epoch. Among the last gomphotheres, it became extinct at the end of the Pleistocene, approximately 12,000 years ago, following the arrival of humans to the Americas.

<i>Stegomastodon</i> Extinct genus of mammals

Stegomastodon is an extinct genus of gomphotheres. It ranged throughout North America from the Pliocene, to the Early Pleistocene. The former South American species have been synonymized with Notiomastodon platensis.

<i>Sinomastodon</i> Extinct genus of gomphothere proboscidean

Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.

<i>Tetralophodon</i> Extinct genus of mammals

Tetralophodon is an extinct genus of "tetralophodont gomphothere" belonging to the superfamily Elephantoidea, known from the Miocene of Afro-Eurasia.

Gnathabelodon is an extinct genus of gomphothere endemic to North America that includes species that lived during the Middle to Late Miocene.

<i>Eubelodon</i> Extinct genus of proboscid

Eubelodon is an extinct genus of gomphothere which lived in North America during the Miocene Epoch. It contains a single species: Eubelodon morrilli.

<i>Zygolophodon</i> Extinct genus of mammutid proboscidean

Zygolophodon is an extinct genus of mammutid proboscidean that lived during the Miocene in Africa, Eurasia, and North America.

<i>Notiomastodon</i> Extinct genus of gomphothere elephantimorph native to South America

Notiomastodon is an extinct genus of gomphothere proboscidean, endemic to South America from the Pleistocene to the beginning of the Holocene. Notiomastodon specimens reached a size similar to that of the modern Asian elephant, with a body mass of 3-4 tonnes. Like other brevirostrine gomphotheres such as Cuvieronius and Stegomastodon, Notiomastodon had a shortened lower jaw and lacked lower tusks, unlike more primitive gomphotheres like Gomphotherium.

<i>Choerolophodon</i> Extinct genus of mammals

Choerolophodon is an extinct genus of proboscidean that lived during the Miocene of Eurasia and Africa. Fossils of Choerolophodon have been found in Africa, Southeast Europe, Turkey, Iraq, Iran, the Indian subcontinent, and China.

<span class="mw-page-title-main">Elephantimorpha</span> Clade of mammals

Elephantimorpha is a clade of proboscideans that contains the Mammutidae (mastodons), as well as Elephantida. All members of this group have the horizontal tooth replacement typical of modern elephants, unlike more primitive members of the Elephantiformes. Like modern elephants, the ancestor of Elephantimorpha was likely capable of communicating via infrasonic calls. While early elephantimorphs generally had lower jaws with an elongated mandibular symphysis at the front of the jaw with well developed lower tusks/incisors, from the Late Miocene onwards, many groups convergently developed brevirostrine (shortened) lower jaws with vestigial or no lower tusks, probably corresponding with the elongation and increasingly dexterity of the trunk allowing it to be used as the primary feeding organ.

<i>Konobelodon</i> Extinct genus of mammals

Konobelodon is an extinct genus of amebelodont proboscidean from the Miocene of Africa, Eurasia and North America.

Eurybelodon is an extinct genus of proboscidean in the family Amebelodontidae. It lived in the Clarendonian age of the Miocene.

Research history of <i>Mammut</i> Studies of an extinct genus of proboscidean

The research history of Mammut is extensive given its complicated taxonomic and non-taxonomic histories, with the earliest recorded fossil finds dating back to 1705 in Claverack, New York during the colonial era of what is now the United States of America. Initially thought to belong to biblical antediluvian giants, the fossils were later determined to belong to a proboscidean species as a result of more complete 18th century finds from the locality of Big Bone Lick in what is now Kentucky. The molars were studied by European and American naturalists, who were generally baffled on its lack of analogue to modern elephants, leading to varying hypothesis on the affinities of the teeth. More complete skeletons were found after the independence of the United States colonies from Great Britain within the early 19th century. American historians of the 21st century have made arguments that the early history of M. americanum finds and studies played major roles in shaping American nationalism on the basis of the large sizes and relative completeness of the fossils to disprove the negative theory of social degeneracy in North America.

References

  1. Paleobiology database: Rhynchotherium basic info
  2. Arroyo-Cabrales, J.; Polaco, O. J.; Laurito, C.; Johnson, E.; Alberdi, M. T.; Zamora, A. L. V. (2007). "The proboscideans (Mammalia) from Mesoamerica". Quaternary International. 169–170. Elsevier: 17–23. Bibcode:2007QuInt.169...17A. doi:10.1016/j.quaint.2006.12.017.
  3. FALCONER, H. (1868): Paleontological Memoirs. Volume II: 74 –75; London (Hardwicke).
  4. C. Frick. 1921. Extinct vertebrate faunas of the badlands of Bautista Creek and San Timoteo Canyon, southern California. University of California Publications in Geology 12(5):277-424
  5. FRICK, C. (1933): New Remains of Trilophodont-Tetralophodont mastodons. – Bulletin American Museum of Natural History, 59: 505 – 652.
  6. OSBORN, H. F. (1923): New subfamily, generic and specific stages in the evolution of the proboscidea. – American Museum Novitates, 99: 1– 4.
  7. OSBORN, H. F. (1936): Proboscidea: a monograph of the discovery, evolution, migration, and extinction of the mastodonts and elephants of the world, vol. 1: Moeritherioidea, Deinotherioidea, Mastodontoidea. New York (The American Museum Press).
  8. OLSEN, S. J. (1957): A new beak-jawed mastodont from Florida. – Journal of the Palaeontological Society of India, 2: 131–135.
  9. 1 2 Mothé, Dimila; Ferretti, Marco P.; Avilla, Leonardo S. (12 January 2016). "The Dance of Tusks: Rediscovery of Lower Incisors in the Pan-American Proboscidean Cuvieronius hyodon Revises Incisor Evolution in Elephantimorpha". PLOS ONE. 11 (1): e0147009. Bibcode:2016PLoSO..1147009M. doi: 10.1371/journal.pone.0147009 . PMC   4710528 . PMID   26756209.
  10. 1 2 LUCAS, S.G. & MORGAN, G.S., 2008. Taxonomy of Rhynchotherium (Mammalia, Proboscidea) from the Miocene-Pliocene of North America.- New Mex. Mus. Nat. Hist. Sci. Bull. 44: 71-87.
  11. LUCAS, S. G. (2010): Rhynchotherium Falconer, 1868 (Mammalia; Proboscidea): proposed conservation of usage by designation of Rhynchotherium falconeri OSBORN, 1923 as the type species. – Bulletin of Zoological Nomenclature, 67: 158 –162.
  12. "OPINION 2295 (Case 3515) Rhynchotherium Falconer, 1868 (Mammalia; Proboscidea): usage conserved by designation of Rhynchotherium falconeri Osborn, 1923 as the type species". March 2012.