The Proboscidea are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From the mid-Miocene onwards, most proboscideans were very large. The largest land mammal of all time may have been a proboscidean; Palaeoloxodon namadicus was up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, almost double the weight of some sauropods like Diplodocus carnegii. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
Deinotherium was a large elephant-like proboscidean that appeared in the Middle Miocene and survived until the Early Pleistocene. Although superficially resembling modern elephants, they had notably more flexible necks, limbs adapted to a more cursorial lifestyle as well as tusks that curved downwards and back. In addition, their tusks didn't emerge from the maxilla as in elephants but from the mandible. Deinotherium was a widespread genus, ranging from East Africa to the south to Europe and east to the Indian Subcontinent. They were browsing animals with a diet mainly consisting of leaves, and they most likely died out as forested areas were gradually replaced by open grassland during the latter half of the Neogene.
Moeritherium is an extinct genus of primitive proboscideans. These prehistoric mammals are related to the elephant and, more distantly, sea cows and hyraxes. They lived during the Eocene epoch.
Arsinoitherium is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, A. giganteum, lived in Ethiopia about 27 million years ago.
Deinotheriidae is a family of prehistoric elephant-like proboscideans that lived during the Cenozoic era, first appearing in Africa, then spreading across southern Asia (Indo-Pakistan) and Europe. During that time, they changed very little, apart from growing much larger in size; by the late Miocene, they had become the largest land animals of their time. Their most distinctive features were the downward-curving tusks on the lower jaw.
Paraceratherium is an extinct genus of hornless rhinoceros. It is one of the largest terrestrial mammals that has existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. It is classified as a member of the family Paraceratheriidae. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species P. bugtiense was originally placed.
Oligopithecus is a fossil primate that lived in Africa during the Early Oligocene. It is represented by one species, Oligopithecus savagei, known from one jaw bone found in Egypt.
The genus Apidium is that of at least three extinct primates living in the early Oligocene, from 30 to 28 million years ago. Apidium fossils are common in the Fayoum deposits of Egypt. Fossils of the earlier species, Apidium moustafai, are rare; fossils of the later species Apidium phiomense are fairly common.
Pyrotherium is an extinct genus of South American ungulate, of the order Pyrotheria, that lived in what is now Argentina and Bolivia, during the Late Oligocene. It was named Pyrotherium because the first specimens were excavated from an ancient volcanic ash deposit. Fossils of the genus have been found in the Deseado and Sarmiento Formations of Argentina and the Salla Formation of Bolivia.
Numidotheriidae is an extinct family of primitive proboscidean that lived from the late Paleocene to the early Oligocene periods of North Africa.
Barytherium is a genus of an extinct family (Barytheriidae) of primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa. The type species is Barytherium grave, found at the beginning of the 20th century in Fayum, Egypt. Since then, more complete specimens have been found at Dor el Talha, Libya. More fossils were also discovered in 2011 in the Aidum area in Dhofar by Oman's Ministry of Heritage and Culture, which was named Barytherium omansi.
Eritreum melakeghebrekristosi is an extinct species of proboscidean mammal, which lived in Northeast Africa during the late Oligocene some 27 million years ago, and is considered to be the missing link between modern elephants and their ancestors. The fossils of this species are the oldest known fossils featuring the horizontal tooth displacement seen in modern elephants. The species is estimated to have weighed 484 kg (1,067 lb) and stood about 1.3 m (4.3 ft) at the shoulder, much smaller than modern species.
Barytheriidae is an extinct family of primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa and Oman. The Barytheriidae were the first large-sized proboscideans to appear in the fossil record and were characterized by a strong sexual dimorphism.
Eogavialis is an extinct genus of eusuchian crocodylomorph, usually regarded as a gavialoid crocodylian. It superficially resembles Tomistoma schlegelii, the extant false gharial, and consequently material from the genus was originally referred to Tomistoma. Indeed, it was not until 1982 that the name Eogavialis was constructed after it was realised that the specimens were from a more basal form.
Astorgosuchus is an extinct monospecific genus of crocodilian, closely related to true crocodiles, that lived in Pakistan during the late Oligocene period. It contains a single species, Astorgosuchus bugtiensis, which was originally named as a species of Crocodylus in 1908 and was moved to its own genus in 2019.
Propliopithecoidea is a superfamily of catarrhine primates that inhabited Africa and Arabia during the Early Oligocene about 32 to 29 million years ago. Fossils have been found in Egypt, Oman and Angola. They are one of the earliest known families of catarrhines.
Dzungariotherium is a genus of indricothere, an extinct group of large, hornless rhinos, which lived during the middle and late Oligocene of northwest China. The type species D. orgosense was described in 1973 based on fossils—mainly teeth—from Dzungaria in Xinjiang, northwest China.
Paulogervaisia is an extinct genus of mammal, belonging to the family Didolodontidae. Its fossilized remains have been found in South America.
Baguatherium is an extinct genus of herbivorous mammal, belonging to the order Pyrotheria. It lived during the Early Oligocene, and its fossilized remains were discovered in Peru.
Dagbatitherium is an extinct genus of proboscideans. So far a single molar from the phosphate basins of Togo in West Africa has been found. The fossil dates to the Middle Eocene, around 47 million years ago. A striking feature of the tooth are the three pairs of cusps oriented transversely to the longitudinal axis of the tooth. This feature is found in more derived proboscideans, which are grouped in the Elephantiformes. For its age, Dagbatitherium is the earliest member of Elephantiformes to date. Furthermore, it is characterized by a low tooth crown and a humped occlusal pattern. The genus was described in 2021.
