Phanagoroloxodon

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Phanagoroloxodon
Temporal range: Late Pliocene–Early Pleistocene
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Phanagoroloxodon.png
Drawing of the skull in various views
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: Elephantidae
Genus: Phanagoroloxodon
Garutt, 1957
Species:
P. mammontoides
Binomial name
Phanagoroloxodon mammontoides
Garutt, 1957

Phanagoroloxodon is a genus of extinct elephant. It is known from one species, Phanagoroloxodon mammontoides, which is described from a partial skull from Russia, of probable Late Pliocene-Early Pleistocene age. [1] [2]

Contents

History of discovery

The holotype of Phanagoroloxodon was found on the banks of the Psekups river in the northwestern Caucasus of Russia, and was given to the Krasnodar State Historical and Archaeological Memorial Museum-Reserve by I.N. Chistyakov in 1885. [2] It was found in the museum's collections by Wadim E. Garutt in 1957, and was named in that same year. [1] [2] Other possible remains of the species include molar teeth described from the nearby Sinyaya Balka site near the eastern shore of the Sea of Azov. [3] In 2005, a second species Phanagoroloxodon irtyshensis was described based on a skull found near Pavlodar in Kazakhstan, but this may represent a specimen of the steppe mammoth (Mammuthus trogontherii). [4]

Description

Skull on display 2023-02-02. Krasnodarskii gosudarstvennyi istoriko-arkheologicheskii muzei DSC 0245.jpg
Skull on display

Phanagoroloxodon resembles Elephas (which contains the living Asian elephant) and mammoths (genus Mammuthus) in many regards. Like Elephas, the top of the skull has a saddle-like groove running along the midline and the nasal process is rounded. On the other hand, the molars have occiput is almost devoid of tubercles, the hind molars lack obliteration figures, the tusks are suggested to be twisted, similar to those of mammoths. [2]

Taxonomy

Phanagoroloxodon has been suggested to be more closely related to Elephas and Mammuthus than to Loxodonta (which contains the living African elephants) due to it combining characteristics of both of these genera, with Garutt proposing that it could be ancestral to both Elephas and Mammuthus. [2] [5] It was assigned to a tribe of its own, Phanagorodontini by Garutt in 1991. [6] A 2020 PhD thesis by Steven Zhang suggested that Elephas recki brumpti from the Pliocene of East Africa should be subsumed into the species Elephas planifrons, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent, and that this species should be placed as a second species of Phanagoroloxodon. [5] However, these suggestions were rejected by Sanders (2023). [7]

Recontruction of Phanagoroloxodon Phanagoroloxodon mammontoides.png
Recontruction of Phanagoroloxodon

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<span class="mw-page-title-main">Proboscidea</span> Order of mammals including elephants

Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.

<span class="mw-page-title-main">Mastodon</span> Extinct genus of proboscideans

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<span class="mw-page-title-main">Elephantidae</span> Family of mammals

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<i>Elephas</i> Genus of mammals

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Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.

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References

  1. 1 2 GARUTT, W.E., 1957. On a new fossil elephant Phanagoroloxodon mammontoides gen. et sp. nov. from the Caucasus. Reports of the Academy of Sciences of the USSR, 112,2: 333-335. (In Russian)
  2. 1 2 3 4 5 W.E. Garutt. (1995). The phanagorian elephant Phanagoroloxodon mammontoides Garutt, 1957 from the Pliocene of the north-western Caucasus. Cranium, 12(2), 87–92.
  3. Baigusheva, Vera S.; Titov, Vadim V.; Foronova, Irina V. (October 2016). "Teeth of early generations of Early Pleistocene elephants (Mammalia, Elephantidae) from Sinyaya Balka/Bogatyri site (Sea of Azov Region, Russia)". Quaternary International. 420: 306–318. doi:10.1016/j.quaint.2015.08.007.
  4. Lister, Adrian M.; Stuart, Anthony J. (December 2010). "The West Runton mammoth (Mammuthus trogontherii) and its evolutionary significance". Quaternary International. 228 (1–2): 180–209. Bibcode:2010QuInt.228..180L. doi:10.1016/j.quaint.2010.07.032.
  5. 1 2 H. Zhang Elephas recki: the wastebasket? 66th Symposium of Vertebrate Palaeontology and Comparative Anatomy, Manchester. (2018)
  6. Garrutt, Wadim E. (1991). "The Phanagorian elephant Phanagoroloxodon mammontoides Garutt, 1957, and the problem of evolutionary ways in the subfamily Elephantinae". The Sixth Coordination Conference for the Study of Mammoths and Mammoth Fauna. Theses of Reports. The Zoological Institute of the Academy of Sciences of the USSR.: 23.
  7. Sanders, William J. (2023-07-07). Evolution and Fossil Record of African Proboscidea (1 ed.). Boca Raton: CRC Press. pp. 267–293. doi:10.1201/b20016. ISBN   978-1-315-11891-8.