Elephas planifrons Temporal range: | |
---|---|
Partial skull in various views | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Proboscidea |
Family: | Elephantidae |
Genus: | Elephas |
Species: | †E. planifrons |
Binomial name | |
†Elephas planifrons Falconer & Cautley, 1846 | |
Elephas planifrons is an extinct species of elephant, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent.
The number of lamellae on the third molars is typically in the range of 11 to 15. [1] Members of the species retained permanent premolars, unlike living Asian elephants. [2]
Isotopic evidence suggests that the species had a grazing based diet. [3]
Elephas planifrons represents one of the earliest dispersals of elephants outside of Africa, first arriving on the Indian subcontinent around 3.6 million years ago [4] [3] (though some authors have suggested a much earlier age over 5 million years ago). The date of the last record of the species is highly uncertain, with estimates ranging from 2.5 million to 800,000 years ago. [5] While only definitvely reported from the Indian subcontinent, possible E. planifrons remains have been reported from Indonesia, and the species may be closely related to the Indonesian dwarf elephant genus Stegoloxodon. [2] A 2020 PhD thesis suggested that the species was not a true member of the genus Elephas , but should instead be considered a member of the genus Phanagoroloxodon with the African Pliocene species "Elephas" recki brumpti considered to be a synonym of "E". planifrons. However, its placement in Phanagoroloxodon and its synonymity with "E". recki brumpti have been rejected by other authors, though the two species may be closely related. [6]
A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene until about 4,000 years ago, with mammoth species at various times inhabiting Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
A mastodon is a member of the genus Mammut, which, strictly defined, was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 27–25 million years ago, during the Oligocene.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene or possibly the late Miocene.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae.
Theropithecus is a genus of primates in the family Cercopithecidae. It contains a single living species, the gelada, native to the Ethiopian Highlands.
Palaeoloxodon recki, often known by the synonym Elephas recki, is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the early Middle to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
Zygolophodon is an extinct genus of mammutid proboscidean that lived during the Miocene in Africa, Eurasia, and North America.
Elephas hysudricus is an extinct elephant species known from the Pleistocene of Asia. It is thought to be ancestral to the living Asian elephant, from which it is distinguished by the molar teeth having a lower crown height and a lower lamellae number. Remains of the species are primarily known from the Indian subcontinent, with the most important remains coming from the Siwalik Hills. The oldest remains of the species in the Siwaliks are placed at around 2.6 million year ago at the beginning of the Early Pleistocene, with the youngest dates in the Siwaliks during the Middle Pleistocene around 0.6 million years ago, though it likely persisted on the subcontinent later than this based on remains found elsewhere.
Palaeoloxodon naumanni is an extinct species of elephant belonging to the genus Palaeoloxodon that was native to the Japanese archipelago during the Middle to Late Pleistocene around 330,000 to 24,000 years ago. It is named after the German geologist Heinrich Edmund Naumann who first described remains of the species in the 19th century, with the species sometimes being called Naumann's elephant. Fossils attributed to P. naumanni are also known from China, though the status of these specimens is unresolved, and some authors regard them as belonging to separate species.
Mammuthus rumanus is a species of mammoth that lived during the Pliocene in Eurasia. It the oldest mammoth species known outside of Africa.
Stegoloxodon is an extinct genus of dwarf elephant known from the Early Pleistocene of Indonesia. It contains two species, S. indonesicus from Java, and S. celebensis from Sulawesi. Its relationship with other elephants is uncertain.
Phanagoroloxodon is an extinct genus of elephant. It is known from one species Phanagoroloxodon mammontoides described from a partial skull found on the banks of the Psekups river in the northwestern Caucasus of Russia, of probable Late Pliocene-Early Pleistocene age. Phanagoroloxodon has been suggested to share a close common ancestry with Elephas, as well as mammoths, combining characteristics of both genera. Like the Asian elephant, the top of the skull has a groove running along the midline, while the tusks are suggested to be twisted, similar to those of mammoths. A 2020 PhD thesis by Steven Zhang suggested that Elephas recki brumpti from the Pliocene of East Africa should be subsumed into the species Elephas planifrons, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent, and that this species should be placed as a second species of Phanagoroloxodon. However, these suggestions were rejected by Sanders (2023).