| Mammuthus rumanus Temporal range: Late Pliocene, | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Proboscidea |
| Family: | Elephantidae |
| Genus: | † Mammuthus |
| Species: | †M. rumanus |
| Binomial name | |
| †Mammuthus rumanus Ștefănescu, 1924 | |
Mammuthus rumanus is a species of mammoth that lived during the Pliocene in Eurasia. It the oldest mammoth species known outside of Africa. [1]
Mammuthus rumanus is suggested to have originated in Africa. [1] Material intermediate between African mammoths and Mammuthus rumanus has been reported from Bethlehem in the Levant, dating to sometime in the Late Pliocene, around 3-4 million years ago. [2] The oldest calibrated dates for Mammuthus rumanus and mammoths outside Africa are from Romania, dating to around 3.2 million years ago. [3] Remains have been reported spanning from Britain to China. [1] It is probably ancestral to Mammuthus meridionalis. [4]
Mammuthus rumanus is only known from fragmentary remains, typically isolated teeth, with a mandible also known. The number of plates on the third molar teeth is around 8-10, consistently lower than is known in other non-African mammoth species, including M. meridionalis. [4]
Studies of specimens from Britain found that they likely consumed browse and inhabited open environments. [5] [6] In Europe, it coexisted alongside other proboscideans, including the mastodon species "Mammut" borsoni and the "tetralophodont gomphothere" Anancus arvernensis, likely niche partitioning with the latter by occupying different habitats. [5] [6]
A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are more closely related to living Asian elephants than African elephants.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Proboscideans include some of the largest known land mammals. The largest land mammal of all time may have been a proboscidean; the elephant Palaeoloxodon namadicus has been estimated to be up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, surpassing the paraceratheres, the otherwise largest known land mammals, though this estimate was made based on a single fragmentary femur and is speculative. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene era.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon remained widespread in South, Southeast and East Asia until the end of the Pleistocene.
Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.
The Columbian mammoth is an extinct species of mammoth that inhabited the Americas from southern Canada to Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian mammoths that colonised North America during the Early Pleistocene around 1.5–1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
The Chibanian, widely known as the Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. The Chibanian name was officially ratified in January 2020. It is currently estimated to span the time between 0.770 Ma and 0.126 Ma, also expressed as 770–126 ka. It includes the transition in palaeoanthropology from the Lower to the Middle Paleolithic over 300 ka.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. It was larger than any living elephant, with adult males suggested to reach 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Cuvieronius is an extinct New World genus of gomphothere which ranged from southern North America to western South America during the Pleistocene epoch. Among the last gomphotheres, it became extinct at the end of the Pleistocene, approximately 12,000 years ago, following the arrival of humans to the Americas.
Stegomastodon is an extinct genus of gomphotheres. It ranged throughout North America from the Pliocene, to the Early Pleistocene. The former South American species have been synonymized with Notiomastodon platensis.
Mammuthus meridionalis, sometimes called the southern mammoth, is an extinct species of mammoth native to Eurasia, including Europe, during the Early Pleistocene, living from around 2.5 million years ago to 800,000 years ago.
Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Primelephas is a genus of Elephantinae that existed during the Miocene and Pliocene epochs. The name of the genus suggests 'first elephant'. These primitive elephantids are thought to be the common ancestor of Mammuthus, the mammoths, and the closely allied genera Elephas and Loxodonta, the Asian and African elephants, diverging some 4-6 million years ago. It had four tusks, which is a trait not shared with its descendants, but common in earlier proboscideans. The type species, Primelephas gomphotheroides, was described by Vincent Maglio in 1970, with the specific epithet indicating the fossil specimens were gomphothere-like. Primelephas korotorensis is the only other species to be assigned to the genus. All fossils found of the Primelephas have been found in Africa, primarily in modern day Chad, Tanzania, Kenya, Ethiopia, and Uganda.
Stegodontidae is an extinct family of proboscideans from Africa and Asia from the Early Miocene to the Late Pleistocene. It contains two genera, the earlier Stegolophodon, known from the Miocene of Asia and the later Stegodon, from the Late Miocene to Late Pleistocene of Africa and Asia which is thought to have evolved from the former. The group is noted for their plate-like lophs on their teeth, which are similar to elephants and different from those of other extinct proboscideans like gomphotheres and mammutids, with both groups having a proal jaw movement utilizing forward strokes of the lower jaw. These similarities with modern elephants were probably convergently evolved. Like elephantids, stegodontids are thought to have evolved from gomphothere ancestors.
Elephas planifrons is an extinct species of elephant, known from the Late Pliocene-Early Pleistocene of the Indian subcontinent.
"Mammut" borsoni is an extinct species of mammutid proboscidean known from the Late Miocene to Early Pleistocene of Eurasia, spanning from western Europe to China. It is the last known mammutid in Eurasia, and amongst the largest of all proboscideans.
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