Pant-y-Ffynnon Quarry fissure fills | |
---|---|
Stratigraphic range: Rhaetian, ~ | |
Type | Fissure fill |
Overlies | Friars Point Limestone Formation |
Lithology | |
Primary | Breccia |
Location | |
Coordinates | 51°27′29″N3°22′26″W / 51.458°N 03.374°W |
Region | South Wales |
Country | Wales |
Pant-y-Ffynnon Quarry is a stone quarry in the Vale of Glamorgan, Wales, around 3 kilometers east of Cowbridge. [1] It contains fissure fill deposits dating to the Late Triassic (Rhaetian), hosted within karsts of Carboniferous aged limestone, primarily the Friars Point Limestone Formation. Remains of numerous small vertebrates, notably archosaurs, are known from the fissure fills in the quarry, similar to other Late Triassic-Early Jurassic fissure fill deposits known from Southwest England and southern Wales.
The quarry was likely in use since at least the 1910s, [2] and the first fossil specimens discovered at the quarry were collected by palaeontologists Kenneth Kermack and Pamela Robinson of University College London between 1951 and 1952, and were first presented at a talk in 1953 and later written on briefly in 1956. [3]
The quarry was abandoned during the 1960s and few fossil discoveries have been made there since. [2] Much of the remains discovered between 1951-52 were left undescribed until the 21st century, with most only initially being briefly described upon discovery.
Taxon | Species | Presence | Notes | Images |
---|---|---|---|---|
Aenigmaspina [2] | A. pantyffynnonensis [2] | Geographically present in Pantyffynnon [4] | Remains include osteoderms, vertebrae, ribs and a scapula clustered tightly together, as well as more pieces from the forelimb and additional vertebrae that likely belong to this individual. [2] Nicknamed 'Edgar'. [3] | |
Clevosaurus [5] | C. cambrica, [5] C. sp. [5] | Geographically present in Pantyffynnon [4] | Partial cranial and post-cranial remains from several specimens, with bite marks belonging to Terrestrisuchus present. [5] | |
Crinoidea [5] | Indeterminate [5] | Geographically present in Pantyffynnon [4] | Reworked from older Carboniferous sediments. [5] | |
Crocodylomorpha [6] | Indeterminate [6] | Geographically present in Pantyffynnon [4] | Isolated bones associated with the holotype of Pendraig . [7] Likely the same animal as Terrestrisuchus | |
Diphydontosaurus [5] | D. sp. [5] | Geographically present in Pantyffynnon [4] | Left dentary. [5] | |
Kuehneosaurus [8] | K. sp. [8] | Geographically present in Pantyffynnon [4] | Indeterminate remains, [9] with Keeble et al. (2018) unable to source this specimen. [5] | |
Lepidosauria | Indeterminate | Geographically present in Pantyffynnon [4] | Indeterminate remains. | |
Pantydraco [10] | P. caducus [10] | Geographically present in Pantyffynnon [4] | Remains include a skull, a partial jawbone, and vertebrae of the cervix, an incomplete right pelvic bone, and partial forelimbs. | |
Pendraig [7] | P. milnerae [7] | Geographically present in Pantyffynnon [4] | Remains include a partial pelvis, vertebrae and a left femur that is from the same individual was found disarticulated from the main block. [7] | |
Rhynchocephalia [5] | Indeterminate [5] | Geographically present in Pantyffynnon [4] | Represented by one or two indeterminate species. [5] | |
Terrestrisuchus [8] | T. gracilis, [8] T. sp.? | Geographically present in Pantyffynnon [4] | Several well-preserved articulated partial skeletons and various isolated bones. [8] | |
Morganucodon is an early mammaliaform genus that lived from the Late Triassic to the Middle Jurassic. It first appeared about 205 million years ago. Unlike many other early mammaliaforms, Morganucodon is well represented by abundant and well preserved material. Most of this comes from Glamorgan in Wales, but fossils have also been found in Yunnan Province in China and various parts of Europe and North America. Some closely related animals (Megazostrodon) are known from exquisite fossils from South Africa.
Archosauria or archosaurs is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistic sense of the term includes all living and extinct relatives of birds and crocodilians such as non-avian dinosaurs, pterosaurs, phytosaurs, aetosaurs and rauisuchians as well as many Mesozoic marine reptiles. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants.
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.
Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution
Thecodontosaurus is a genus of herbivorous basal sauropodomorph dinosaur that lived during the late Triassic period.
Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.
Terrestrisuchus is an extinct genus of very small early crocodylomorph that was about 76 centimetres (30 in) long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by type and only known species T. gracilis. Terrestrisuchus was a long-legged, active predator that lived entirely on land, unlike modern crocodilians. It inhabited a chain of tropical, low-lying islands that made up southern Britain, along with similarly small-sized dinosaurs and abundant rhynchocephalians. Numerous fossils of Terrestrisuchus are known from fissures in limestone karst which made up the islands it lived on, which formed caverns and sinkholes that preserved the remains of Terrestrisuchus and other island-living reptiles.
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably more ecologically diverse than modern crocodillians. The earliest and most primitive crocodylomorphs are represented by "sphenosuchians", a paraphyletic assemblage containing small-bodied, slender forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, crocodylomorphs morphologically diversified into numerous niches, with the subgroups Neosuchia and the extinct Thalattosuchia adapting to aquatic life, while some terrestrial groups adopted herbivorous and omnivorous lifestyles. Terrestrial crocodylomorphs would continue to co-exist alongside aquatic forms until becoming extinct during the Miocene.
Trilophosaurus is a lizard-like trilophosaurid allokotosaur known from the Late Triassic of North America. It was a herbivore up to 2.5 m long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak.
Pantydraco is a genus of basal sauropodomorph dinosaur from the Late Triassic of the United Kingdom (Wales). It is based on a partial juvenile skeleton once thought to belong to Thecodontosaurus. Only one valid species of Pantydraco is recognised: P. caducus.
Kuehneotherium is an early mammaliaform genus, previously considered a holothere, that lived during the Late Triassic-Early Jurassic Epochs and is characterized by reversed-triangle pattern of molar cusps. Although many fossils have been found, the fossils are limited to teeth, dental fragments, and mandible fragments. The genus includes Kuehneotherium praecursoris and all related species. It was first named and described by Doris M. Kermack, K. A. Kermack, and Frances Mussett in November 1967. The family Kuehneotheriidae and the genus Kuehneotherium were created to house the single species Kuehneotherium praecursoris. Modeling based upon a comparison of the Kuehneotherium jaw with other mammaliaforms indicates it was about the size of a modern-day shrew between 4 and 5.5 g at adulthood.
Clevosaurus is an extinct genus of rhynchocephalian reptile from the Late Triassic and the Early Jurassic periods. Species of Clevosaurus were widespread across Pangaea, and have been found on all continents except Australia and Antarctica. Five species of Clevosaurus have been found in ancient fissure fill deposits in south-west England and Wales, alongside other sphenodontians, early mammals and dinosaurs. In regards to its Pangaean distribution, C. hadroprodon is the oldest record of a sphenodontian from Gondwana, though its affinity to Clevosaurus has been questioned.
The Magnesian Conglomerate is a geological formation in Clifton, Bristol in England, Gloucestershire and southern Wales, present in Tytherington, Durdham Down, Slickstones Quarry and Cromhall Quarry.
Heptasuchus is an extinct genus of loricatan pseudosuchian known from the Middle or Late Triassic upper Chugwater Group of Wyoming, United States. It contains a single species, Heptasuchus clarki, the first formally recognized "rauisuchian" or loricatan pseudosuchian from North America.
Tricuspisaurus is an extinct genus of reptile originally described as a trilophosaurid; it was later considered likely to be a procolophonid, but recent analyses have affirmed the original classification. Fossils are known from the Ruthin Quarry in Glamorgan, Wales, one of several Late Triassic to Early Jurassic British fissure deposits. Like some trilophosaurs, it has an edentulous, or toothless beak. Tricuspisaurus gets its name from its heterodont dentition, which includes tricuspid teeth, or teeth with three cusps. The type species, T. thomasi, was named in 1957 along with the possible trilophosaur Variodens inopinatus from Somerset, England.
Bridetherium is an extinct genus of morganucodontan from Early Jurassic deposits of southern Wales, United Kingdom. Bridetherium is known from some isolated upper and lower molariform. It was collected in the Pant Quarry, Vale of Glamorgan. It was first named by William A. Clemens Jr. in 2011 and the type species is Bridetherium dorisae. The species name is in honour of early mammal paleontologist Doris Mary Kermack.
Aenigmaspina is an extinct genus of enigmatic pseudosuchian (=crurotarsan) archosaur from the Late Triassic of the United Kingdom. Its fossils are known from the Pant-y-ffynnon Quarry in South Wales, of which its type and only known species is named after, A. pantyffynnonensis. Aenigmaspina is characterised by the unusual spines on its vertebrae, which are broad and flat on top with a unique 'Y' shape. Although parts of its skeleton is relatively well known, the affinities of Aenigmaspina to other pseudosuchians are unclear, although it is possibly related to families Ornithosuchidae, Erpetosuchidae or Gracilisuchidae.
The Avon Fissure Fill, also known as the Bristol Fissure Fill or Tytherington Fissure Fill, is a fissure fill in Avon, England which dates variously from the Norian and Rhaetian stages of the Late Triassic, or possibly as late as the Hettangian stage of the Early Jurassic. The fissure fill at Avon was a sinkhole formed by the dissolution of Lower Carboniferous limestones.
Pendraig is a genus of coelophysoid theropod dinosaur from South Wales. It contains one species, Pendraig milnerae, named after Angela Milner. The specimen was discovered in the Pant-y-Ffynnon quarry. In life it would have measured 1 m (3.3 ft) in length.
Palaeontology in Wales is palaeontological research occurring in Wales.
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