Petalocrinidae

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Petalocrinidae
Temporal range: Early Ordovician–Early Devonian
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Petalocrinidae

Weller et Davidson, 1896
Genera

Petalocrinidae is an extinct family of cladid crinoids from the Early Ordovician to Lower Devonian. Fossils of petalocrinoids have been found in China, Europe and the United States.

Cladida order of echinoderms

Cladida is an extinct order of crinoids from the Middle to Late Devonian.

Crinoid Class of echinoderms

Crinoids are marine animals that make up the class Crinoidea, one of the classes of the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. Those crinoids which, in their adult form, are attached to the sea bottom by a stalk are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, being members of the largest crinoid order, Comatulida.

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

Contents

All petalocrinoids have five, flattened arms formed through the fusion of the branches of the ancestral crinoid arm, an Ordovician crinoid possibly similar to the Silurian Gissocrinus or Arachnocrinus . [1] The shape of these fused arms varies from genus to genus, and from species to species. The upper surface of the arms had distinctive patterns of grooves which are thought to have born captured food particles towards the mouth. The purpose of the arms, themselves, remain unknown, though, some researchers suspect that they may deterred predators by presenting a would-be attacker with a largely inedible chunk of calcite.

Genera

Eopetalocrinus

Eopetalocrinus is the earliest known petalocrinoid, known from a single set of five arms from Arenig-aged strata of Southern China.

Arenig group

In geology, the Arenigian refers both to a time interval during the Lower Ordovician period and also to the suite of rocks which were deposited during this interval.

Petalocrinus

Petalocrinus is the most successful genus in the family, with numerous species found in Lower Silurian strata in the United States, England, Gotland, and China. The arms of Petalocrinus are usually wedge-shaped, akin to a guitar pick, though, some species, like the English P. bifurcatus, have their own distinct shapes (in the case of P. bifurcatus, each arm is bifurcated into the shape of a "V").

United States Federal republic in North America

The United States of America (USA), commonly known as the United States or America, is a country comprising 50 states, a federal district, five major self-governing territories, and various possessions. At 3.8 million square miles, the United States is the world's third or fourth largest country by total area and is slightly smaller than the entire continent of Europe's 3.9 million square miles. With a population of over 327 million people, the U.S. is the third most populous country. The capital is Washington, D.C., and the most populous city is New York City. Forty-eight states and the capital's federal district are contiguous in North America between Canada and Mexico. The State of Alaska is in the northwest corner of North America, bordered by Canada to the east and across the Bering Strait from Russia to the west. The State of Hawaii is an archipelago in the mid-Pacific Ocean. The U.S. territories are scattered about the Pacific Ocean and the Caribbean Sea, stretching across nine official time zones. The extremely diverse geography, climate, and wildlife of the United States make it one of the world's 17 megadiverse countries.

England Country in north-west Europe, part of the United Kingdom

England is a country that is part of the United Kingdom. It shares land borders with Wales to the west and Scotland to the north. The Irish Sea lies west of England and the Celtic Sea to the southwest. England is separated from continental Europe by the North Sea to the east and the English Channel to the south. The country covers five-eighths of the island of Great Britain, which lies in the North Atlantic, and includes over 100 smaller islands, such as the Isles of Scilly and the Isle of Wight.

Gotland Island and historical province in Sweden

Gotland is a province, county, municipality, and diocese of Sweden. It is Sweden's largest island. The province includes the islands of Fårö and Gotska Sandön to the north, as well as the Karlsö Islands to the west. The population is 58,595, of which about 23,600 live in Visby, the main town. The island of Gotland and the other areas of the province of Gotland make up less than one percent of Sweden's total land area.

Sinopetalocrinus

Sinopetalocrinus is one of three petalocrinoid genera endemic to Chinese strata, and is thought to be the daughter genus of Petalocrinus, the result of a diversification event in the early Silurian that produced it, and Spirocrinus. Sinopetalocrinus is very similar in anatomy to Petalocrinus.

In the geological timescale, the Llandovery epoch occurred at the beginning of the Silurian period. The Llandoverian epoch follows the massive Ordovician-Silurian extinction events, which led to a large decrease in biodiversity and an opening up of ecosystems.

Spirocrinus

Spirocrinus is one of three petalocrinoid genera endemic to Chinese strata. It is found in Lower Silurian strata together with species of its sister genus Sinopetalocrinus, and parent genus Petalocrinus. Spirocrinus is unmistakably distinct, as its arms are modified into cigar-shapes, covered in feeding grooves.

Vadarocrinus

Vadarocrinus is the only petalocrinoid genus known from the Devonian, being found in lower Devonian strata of the Koněprusy Limestone, Bohemia. The arms are curved downward. Because the paleoenvironment is thought to have been filled with turbulent water currents, and that a recently found stem ossicle suggests a thick stem, Vadarocrinus is thought to have had a very short stem, compared to other petalocrinoids. [2]

Bohemia Historical land in Czech Republic

Bohemia is the westernmost and largest historical region of the Czech lands in the present-day Czech Republic. In a broader meaning, Bohemia sometimes refers to the entire Czech territory, including Moravia and Czech Silesia, especially in a historical context, such as the Lands of the Bohemian Crown ruled by Bohemian kings.

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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

Llano Uplift

The Llano uplift is a low geologic dome that is about 90 miles (140 km) in diameter. It consists of an island-like exposure of Precambrian igneous and metamorphic rocks surrounded by outcrops of Paleozoic and Cretaceous sedimentary strata. At their widest, the exposed Precambrian rocks extend about 65 miles (105 km) westward from the valley of the Colorado River and beneath a broad, gentle topographic basin drained by the Llano River. The subdued topographic basin is underlain by Precambrian rocks and bordered by a discontinuous rim of flat-topped hills. These hills are the dissected edge of the Edwards Plateau, which consist of overlying Cretaceous sedimentary strata. Within this basin and along its margin are down-faulted blocks and erosional remnants of Paleozoic strata which form prominent hills.

Protaspididae family of fishes (fossil)

Protaspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protaspidids were once thought to represent a transitional form between the Pteraspididae and the Psammosteida, bearing the broad head shield shape of the latter, due to a more benthic (bottom-dwelling) existence, but recent phylogenical comparisons demonstrate that the protaspidids are actually highly derived pteraspidids, and that the anchipteraspidids, the most primitive of pteraspidids, are the sister-group of the Psammosteids.

Cheloniellida is an unranked taxon of extinct Paleozoic arthropods, currently thought to be a member of the Arachnomorpha and related to the Chelicerata and the Trilobita, while not included in any of those. There are eight included genera, whose fossils are found in marine strata ranging from Ordovician to Devonian in age.

Dunkleosteidae family of fishes (fossil)

Dunkleosteidae is an extinct family of arthrodire placoderms. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group. While they were previously thought to be close relatives of the genus Dinichthys and grouped together in the family Dinichthyidae, more recent studies have shown that the two taxa represent two very distinct clades within Arthrodira. The reappraisal of Kiangyousteus lead to a restructuring of the family, with the inclusions of the benthic, aberrant Heterosteus as the sister taxon of Dunkleosteus, and the Late Emsian Xiangshuiosteus as the sister taxon of Eastmanosteus calliaspis, and the removal of Westralichthys from the family

<i>Lunaspis</i> genus of fishes (fossil)

Lunaspis is an extinct genus of armor-plated petalichthyid placoderm fish that lived in shallow marine environments of the Early Devonian period, from approximately 409.1 to 402.5 million year ago. Fossils have been found in Germany, China and Australia. There are three different identified species of within the genus Lunaspis: L. broilii, L. heroldi, and L. prumiensis.

<i>Drepanopterus</i>

Drepanopterus is an extinct genus of eurypterid and the only member of the family Drepanopteridae within the Mycteropoidea superfamily. There are currently three species assigned to the genus. The genus has historically included more species, with nine species associated with the genus Drepanopterus, however five of these have since been proven to be synonyms of pre-existing species, assigned to their own genera, or found to be based on insubstantial fossil data. The holotype of one species proved to be a lithic clast.

Michelinoceras is the oldest known genus of the Michelinocerida, more commonly known as the Orthocerida, characterized by long, slender, nearly cylindrical orthocones with a circular cross section, long camerae, very long body chambers, and a central or near central tubular siphuncle free of organic deposits. Septal necks are straight; connecting rings cylindrical and thin. Cameral deposits are well developed. A radula has been found in one species, with seven teeth per row. It had ten arms, two of which formed longer tentacles.

The Actinocerida comprise an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about 60 to 90 cm but some, like the Huroniidae of the Silurian grew significantly larger.

Junggaria was a genus of rhyniophyte-like land plants known from fossils found in China in Upper Silurian strata. It bore leafless dichotomously or pseudomonopodially branching axes, some of which ended in spore-forming organs or sporangia of complex shape. The genus Cooksonella, found in Kazakhstan from deposits of a similar age, is considered to be an illegitimate synonym.

Yarravia is a genus of extinct vascular plants mainly known from fossils found in Victoria, Australia. Originally the rocks in which they were found were considered to be late Silurian in age; more recently they have been found to be Early Devonian. Specimens consist only of incomplete leafless stems, some of which bore groups of spore-forming organs or sporangia which were fused, at least at the base.

Fossils of many types of water-dwelling animals from the Devonian period are found in deposits in the U.S. state of Michigan. Among the more commonly occurring specimens are bryozoans, corals, crinoids, and brachiopods. Also found, but not so commonly, are armored fish called placoderms, snails, sharks, stromatolites, trilobites and blastoids.

<i>Strophomena</i> genus of brachiopods (fossil)

Strophomena is a genus of brachiopods belonging to the order Strophomenida family Strophomenidae, named by Rafinesque in 1824. They were stationary epifaunal suspension feeders.

Pteraspidiformes order of fishes (fossil)

Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada.

Protopteraspididae family of fishes (fossil)

Protopteraspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protopteraspidids were once thought to represent a taxon of basal pteraspidids but recent evaluations demonstrate that Protopteraspididae is a paraphyletic group of various transitional forms representing a gradual transition between the more advanced Pteraspoidei, and the anchipteraspidids and the Psammosteids.

Entomaspididae family of arthropods (fossil)

Entomaspididae is a family of harpetid trilobites that ranges from the Upper Cambrian to Lower Ordovician of marine strata in China and the United States.

<i>Odontopleura</i>

Odontopleura is a genus of spinose odontopleurid trilobite in the family Odontopleuridae, and is the type genus of that family and of Odontopleurida. The various species are found in Upper Ordovician to Middle Devonian marine strata throughout the world. The best studied fossils are of the type species, O. ovata, from the Wenlock-aged Liteň Formation in Loděnice, in Bohemia, Czech Republic, and, southeastern Gotland, of Sweden.

Diploporita is an extinct class of blastozoan that ranged from the Ordovician to the Devonian. These echinoderms are identified by a specialized respiratory structure, called diplopores. Diplopores are a double pore system that sit within a depression on a single thecal (body) plate; each plate can contain numerous diplopore pairs.

<i>Eusarcana</i> Extinct genus of sea scorpions

Eusarcana is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Eusarcana have been discovered in deposits ranging in age from the Early Silurian to the Early Devonian. Classified as part of the family Carcinosomatidae, the genus contains three species, E. acrocephalus, E. obesus and E. scorpionis, from the Silurian-Devonian of Scotland, the Czech Republic and the United States respectively.

References

  1. Bather, F. A. (1898). Petalocrinus, Weller & Davidson. Harvard University. p. 437.
  2. Prokop, Rudolf J., and Václav Petr. "New finds of Vadarocrinus vassa Prokop, 1984 (Crinoidea, Petalocrinidae) in the Konĕprusy and Lodĕnice Limestones (Lower Devonian, Pragian) of the Barrandian area, Czech Republic." Journal of the National Museum (Prague) National History Series 179 (2010).