Cladida

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Cladida
Temporal range: Ordovician–recent
Ulrichicrinus coryphaeus.jpg
Ulrichicrinus coryphaeus , a eucladid from the Mississippian-age Edwardsville Formation
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Echinodermata
Class: Crinoidea
Subclass: Pentacrinoidea
Infraclass: Inadunata
Parvclass: Cladida
Moore and Laudon (1943) [1]
Subgroups

Cladida is a major subgroup of crinoids with a complicated taxonomic history. Cladida was originally applied to a wide assortment of extinct crinoids with a dicyclic calyx. Under this original definition, cladids would represent a paraphyletic order ancestral to several other major crinoid groups, particularly the living Articulata. More recently, Cladida has been redefined as a monophyletic parvclass of pentacrinoids which encompasses articulates and the extinct Flexibilia (flexibles). Cladids also include various minor taxa such as the hybocrinids and "cyathocrines". As flexibles were not originally considered cladids, the new subgroup Eucladida has been erected for cladids which are more derived than flexibles. Cladida is the sister group to Disparida, another large group of extinct crinoids. [2]

Contents

Taxonomy

List of genera

Related Research Articles

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Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their juvenile form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths as great as 9,000 meters (30,000 ft).

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<span class="mw-page-title-main">Caenogastropoda</span> Clade of sea snails

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<span class="mw-page-title-main">Batrachomorpha</span> Clade of amphibians

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<span class="mw-page-title-main">Laurasiatheria</span> Clade of mammals

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<span class="mw-page-title-main">Eumalacostraca</span> Subclass of crustaceans

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<span class="mw-page-title-main">Articulata (Crinoidea)</span> Subclass of crinoids

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<span class="mw-page-title-main">Taxonomic rank</span> Level in a taxonomic hierarchy

In biology, taxonomic rank is the relative level of a group of organisms in an ancestral or hereditary hierarchy. A common system of biological classification (taxonomy) consists of species, genus, family, order, class, phylum, kingdom, and domain. While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based on genetic analysis have opened the road to cladistics.

<span class="mw-page-title-main">Orthoceratoidea</span> Extinct subclass of cephalopods

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<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

<span class="mw-page-title-main">Bourgueticrinida</span> Extinct order of crinoids

Bourgueticrinida is an order of crinoids that typically live deep in the ocean. Members of this order are attached to the seabed by a slender stalk and are known as sea lilies. While other groups of crinoids flourished during the Permian, bourgueticrinids along with other extant orders did not appear until the Triassic, following a mass extinction event in which nearly all crinoids died out.

<span class="mw-page-title-main">Camerata (Crinoidea)</span> Extinct subclass of crinoids

The Camerata or camerate crinoids are an extinct subclass of Paleozoic stalked crinoids. They were some of the earliest crinoids to originate during the Early Ordovician, reached their maximum diversity during the Mississippian, and became extinct during the Permian–Triassic extinction event. Camerates are the sister group of Pentacrinoidea, which contains all other crinoids. The two largest camerate subgroups are the orders Diplobathrida and Monobathrida.

Disparida is an parvclass of extinct marine animals in the class Crinoidea. Disparids are a speciose and morphologically diverse group of crinoids distinguished by their monocyclic calyx and slender arms without pinnules. They range from the Early Ordovician (Tremadocian) to Middle Permian, reaching their highest diversity during the Late Ordovician.

Pentacrinoidea is a subclass of crinoids containing all members of Crinoidea except for the exclusively Paleozoic camerates. It was originally named in 1918 by Otto Jaekel, who hypothesized a fundamental split between camerate and non-camerate crinoids. Later workers doubted this interpretation, and Pentacrinoidea was rarely used during the rest of the 20th century. Recent phylogenetic work has provided strong support for Jaekel's hypothesis, and Pentacrinoidea was reinstated in a 2017 revision of crinoid systematics.

<span class="mw-page-title-main">Flexibilia</span> Superorder of crinoids

Flexibilia is a superorder of specialized Paleozoic crinoids. They exhibited a conserved body plan and consistent suite of characteristics throughout their long history. Previously considered a subclass with unclear affinities, later investigation determined that flexibles are well-nested within Cladida, a broad group ancestral to living articulate crinoids. The Ordovician cladid Cupulocrinus acts as an intermediate form linking the generalized anatomy of other early cladids with the distinctive anatomy of flexibles, and several studies have considered it to be ancestral to the rest of the group. Although flexibles never reached the same abundance or diversity as many other crinoid groups, they remained a reliable component of crinoid faunas, particularly from the Silurian onwards. Flexible fossils are very rare in the Ordovician, but the Late Ordovician appears to have been an interval of rapid diversification for the group.

References

  1. Evolution and Classification of Paleozoic Crinoids, Raymond Cecil Moore, Lowell Robert Laudon, 1943
  2. 1 2 3 4 Wright, David F.; Ausich, William I.; Cole, Selina R.; Peter, Mark E.; Rhenberg, Elizabeth C. (2017). "Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata)". Journal of Paleontology. 91 (4): 829–846. doi: 10.1017/jpa.2016.142 . ISSN   0022-3360.
  3. 1 2 3 4 5 Wright, David F. (2017-10-23). "Phenotypic Innovation and Adaptive Constraints in the Evolutionary Radiation of Palaeozoic Crinoids". Scientific Reports. 7 (1): 13745. doi: 10.1038/s41598-017-13979-9 . ISSN   2045-2322.