| Pustulinae Temporal range: | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Brachiopoda |
| Class: | † Strophomenata |
| Order: | † Productida |
| Family: | † Echinoconchidae |
| Subfamily: | † Pustulinae Waterhouse, 1981 |
| Subgroups | |
Pustulinae is an extinct subfamily of medium to large-sized brachiopods which lived during the Early Carboniferous period in marine habitats. It is named after the type genus Pustula. [1]
The exact evolutionary relationships of Pustulinae relative to other groups of the suborder Productidina have been a matter of extensive debate throughout much of the 20th Century, primarily due to the three genera Buxtonia , Pustula and Juresania (with the debate later expanded to their families and subfamilies) shifting in position repeatedly between phylogenies and classifications. The emphasis on internal versus external characters to determine the systematics of these groups has largely been responsible for this: the original Treatise on Invertebrate Paleontology published in 1965 placed emphasis on internal characters (including the cardinal process) as diagnostic, whereas the 2000 revision primarily used external features and shell shape, resulting in differing classification of these clades. [1]
More recently, Leighton & Maples (2002) conducted multiple phylogenetic analyses which are strongly in agreement that the four subfamilies Buxtoniinae, Echinoconchinae, Pustulinae and Juresaniinae form the family Echinoconchidae. The results of their phylogenetic analyses are displayed in the cladogram below: [2]
Nektaspida is an extinct order of non-mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous. They were most common however from the Ordovician to the Devonian.
The Treatise on Invertebrate Paleontology published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, taxa with no fossil record whatsoever have just a very brief listing.
Prolecanitoidea is a taxonomic superfamily of ammonoids in the order Prolecanitida. Prolecanitoidea is one of two superfamilies in the order, along with the younger and more complex Medlicottioidea. The Prolecanitoidea were a low-diversity and morphologically conservative group. They lived from the Lower Carboniferous up to the Middle Permian. Their shells are generally smooth and discoidal, with a rounded lower edge, a moderate to large umbilicus, and goniatitic to ceratitic sutures. Suture complexity varies from 10 up to 22 total lobes ; new lobes are added from subdivision of saddles adjacent to the original main umbilical lobe.
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.
The Stipeae are a tribe of grasses within the subfamily Pooidae, with up to 600 described species.
The ”Fallotaspidoidea” are a superfamily of trilobites, a group of extinct marine arthropods. It lived during the Lower Cambrian (Atdabanian) and species occurred on all paleocontinents except for the Gondwana heartland. A member of this group, Profallotaspis jakutensis, has long been the earliest known trilobite, but recently the redlichiid Lemdadella has been claimed as occurring even earlier.
The Camerata or camerate crinoids are an extinct subclass of Paleozoic stalked crinoids. They were some of the earliest crinoids to originate during the Early Ordovician, reached their maximum diversity during the Mississippian, and became extinct during the Permian–Triassic extinction event. Camerates are the sister group of Pentacrinoidea, which contains all other crinoids. The two largest camerate subgroups are the orders Diplobathrida and Monobathrida.
Dictyonellidae is a family of sponges in the order Bubarida, based on the genus DictyonellaSchmidt, 1868.
Diaphragmus is an extinct genus of brachiopod belonging to the order Productida and family Linoproductidae. Specimens have been found in Carboniferous beds in North America.
Pugnoides is an extinct genus of brachiopod belonging to the order Rhynchonellida and family Petasmariidae. Specimens have been found in Devonian to Permian beds in North America, Asia, Europe, western Australia, New Zealand,and New Zealand. The genus was particularly widespread in the Visean.
Productidae is an extinct family of brachiopods which lived from the Upper Devonian to Upper Permian periods in marine environments. It is the most diversified family in the suborder Productidina, with some 100 genera.
Echinoconchidae is an extinct family of brachiopods which lived from the Lower Carboniferous to Upper Permian periods in marine habitats. Currently, four subfamilies are assigned to it, though the evolutionary relationships between them and the family Productidae have been heavily debated for the better part of the 20th Century.
Echinoconchinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats. The family had a cosmopolitan distribution.
Echinoconchus is an extinct genus of brachiopods which lived during the Lower Carboniferous period. The genus was abundant and had a cosmopolitan distribution.
Echinaria is an extinct genus of brachiopods which lived during the Upper Carboniferous and Lower Permian periods. Its fossils have been found in Eurasia, North America and northern South America.
Juresaniinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats.
Antiquatonia is an extinct genus of brachiopod belonging to the order Productida and family Productidae. Specimens have been found in Carboniferous beds across many continents, suggesting the genus had a cosmopolitan distribution. Species level taxonomy of Antiquatonia is in need of revision.
Pustula is an extinct genus of brachiopods which lived during the Carboniferous period. It is the type genus of the subfamily Pustulinae. Its fossils have been found in Eurasia, North America and northern Africa.
Pulchratia is an extinct genus of brachiopods which lived in marine habitats during the Upper Carboniferous period. Its fossils have been found in North America.