The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.
Pareiasaurs are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with osteoderms which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct in the Permian–Triassic extinction event.
The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlies the Stormberg Group. Based on stratigraphic position, lithostratigraphic and biostratigraphic correlations, palynological analyses, and other means of geological dating, the Beaufort Group rocks are considered to range between Middle Permian (Wordian) to Early Triassic (Anisian) in age.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
The Tapinocephalus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the middle Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 2,000 metres (6,600 ft), occur from Merweville and Leeu-Gamka in its southernmost exposures, from Sutherland through to Beaufort West where outcrops start to only be found in the south-east, north of Oudshoorn and Willowmore, reaching up to areas south of Graaff-Reinet. Its northernmost exposures occur around the towns Fraserburg and Victoria West. The Tapinocephalus Assemblage Zone is the second biozone of the Beaufort Group.
The Ecca Group is the second of the main subdivisions of the Karoo Supergroup of geological strata in southern Africa. It mainly follows conformably after the Dwyka Group in some sections, but in some localities overlying unconformably over much older basement rocks. It underlies the Beaufort Group in all known outcrops and exposures. Based on stratigraphic position, lithostratigraphic correlation, palynological analyses, and other means of geological dating, the Ecca Group ranges between Early to earliest Middle Permian in age.
Shihtienfenia was a pareiasaurid parareptile from the Late Permian of China.
The Tambach Formation is an Early Permian-age geologic formation in central Germany. It consists of red to brown-colored sedimentary rocks such as conglomerate, sandstone, and mudstone, and is the oldest portion of the Upper Rotliegend within the Thuringian Forest Basin.
The Ermaying Formation is a geological formation of Anisian age in north-central China. It is found across much of the Ordos Basin, at outcrops within the provinces of Shaanxi, Shanxi, and Inner Mongolia. It is composed of up to 600 m thick sequence of mudstone and sandstone, overlying the Heshanggou Formation and underlying the Tongchuan Formation. In the southern part of the Ordos Basin, the Zhifang Formation is equivalent to the Ermaying Formation.
The Usili Formation is a Late Permian geologic formation in Tanzania. It preserves fossils of many terrestrial vertebrates from the Permian, including temnospondyls, pareiasaurs, therapsids and the archosauromorph Aenigmastropheus.
The Balfour Formation is a geological formation that is found in the Beaufort Group, a major geological group that forms part of the greater Karoo Supergroup in South Africa. The Balfour Formation is the uppermost formation of the Adelaide Subgroup which contains all the Late Permian - Early Triassic aged biozones of the Beaufort Group. Outcrops and exposures of the Balfour Formation are found from east of 24 degrees in the highest mountainous escarpments between Beaufort West and Fraserburg, but most notably in the Winterberg and Sneeuberg mountain ranges near Cradock, the Baviaanskloof river valley, Graaff-Reniet and Nieu Bethesda in the Eastern Cape, and in the southern Free State province.
The Katberg Formation is a geological formation that is found in the Beaufort Group, a major geological group that forms part of the greater Karoo Supergroup in South Africa. The Katberg Formation is the lowermost geological formation of the Tarkastad Subgroup which contains the Lower to Middle Triassic-aged rocks of the Beaufort Group. Outcrops and exposures of the Katberg Formation are found east of 24 degrees on wards and north of Graaff-Reniet, Nieu Bethesda, Cradock, Fort Beaufort, Queensdown, and East London in the south, and ranges as far north as Harrismith in deposits that form a ring around the Drakensberg mountain ranges.
The Teekloof Formation is a geological formation that forms part of the Beaufort Group, one of the five geological groups that comprises the Karoo Supergroup in South Africa. The Teekloof Formation is the uppermost formation of Adelaide Subgroup deposits West of 24ºE and contains Middle to Late Permian-aged deposits and four biozones of the Beaufort Group. It overlies the Abrahamskraal Formation. The Teekloof Formation does not underlie other units other than the younger Karoo dolerites and sills that relate to the emplacement of the Early Jurassic Drakensberg Group to the east. Outcrops and exposures of the Teekloof Formation range from Sutherland through the mountain escarpments between Fraserburg and Beaufort West. The northernmost localities of the Teekloof Formation are found by Loxton, Victoria West and Richmond.
The Naobaogou Formation is a geological formation in the Daqing Mountains of China. It is likely of Lopingian age. It consists of three rhythms of sediment, labelled members I-III primarily of purple siltstone, but each with a thick basal conglomerate bed. It is notable for its fossil content, producing one of the most diverse Late Permian vertebrate faunas outside Russia and South Africa.
The Moradi Formation is a geological formation in Niger. It is of Late Permian age. It is informally divided into three subunits. The lower portion of the formation consists of red mudstone, with muddy calcareous sandstone and quartz-granlule conglomerate present as lenses. The middle portion consists of muddy siltstone in thick beds interbedded with red argillaceous sandstone. The lower two thirds of the upper portion of the formation consist of red siltstone intercalated with channel lag intraformational conglomerates, while the upper third consists of barchanoid shaped lenses of conglomeratic sandstone with ventifacts. These facies are indicatived of deposition under arid conditions, with less than 300 millimetres (12 in) of annual rainfall in the Central Pangean desert, with annual temperatures of 30 to 35 °C, but with ephemeral water presence including lakes.
The Ntawere Formation is a Middle Triassic (Anisian) geological formation in Zambia, preserving fossils of synapsids, archosaurs, and temnospondyls.
Taoheodon is an extinct genus of dicynodont therapsid from the Sunjiagou Formation in the Shanxi province of China, dated to the Wuchiapingian age of the Late Permian. Its type and only known species is T. baizhijuni. Taoheodon was a close relative of the well known Dicynodon, and may represent a biogeographical link between the South African Dicynodon and similar dicynodonts found in Laos.
Aversor is an extinct genus of embolomere which lived in the Early Permian of Russia. It contains a single species, Aversor dmitrievi, which is based on skull and jaw fragments from the Intinskaya Svita near Pechora. It may have been the youngest known eogyrinid, and was the youngest known embolomere until the discovery of Seroherpeton, a Late Permian embolomere described in 2020. Aversor and Seroherpeton both lived at higher latitudes than older embolomeres, suggesting that the group abandoned arid equatorial areas prior to their final demise.
Seroherpeton is an extinct genus of embolomere known from the Late Permian aged Sunjiagou Formation of Northern China. It is the youngest known embolomere, with the previous youngest members being from the Cisuralian, and also lived further north than any other embolomere, with all other known members of the group being from Euramerica. It is known from a partial skull, consisting of portions of the maxilla, ectopterygoid, pterygoid and quadrate.
Sanchuansaurus is an extinct genus of pareiasaur from the late Permian Sunjiagou Formation of China. The genus contains a single species, S. pygmaeus, known from a partial maxilla and left leg bones. A 2013 study suggested that both Sanchuansaurus and another Chinese pareiasaur, Huanghesaurus, were synonymous with Shansisaurus. However, this has not been followed in future research, with later papers considering Sanchuansaurus to be distinct, with Huanghesaurus and Shansisaurus possibly being synonymous with Shihtienfenia, another Chinese pareiasaur.
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