Terrestrial molluscs or land molluscs (mollusks) are an ecological group that includes all molluscs that live on land in contrast to freshwater and marine molluscs. They probably first occurred in the Carboniferous, arising from freshwater ones. [1]
This group includes land snails and land slugs. Loss of the shell has taken place many times in different groups that are not evolutionarily closely related, and land snails and slugs are most often treated together as a single group in specialized malacological literature. [2] [3]
All terrestrial molluscs belong to the class Gastropoda. However, colonization of the land took place several times during the evolutionary past, and as a result terrestrial molluscs are classified in several different, often not closely related, gastropod taxa. [2]
Terrestrial mollusks comprise about 35 thousand species, most of which belong to the order (in some sources suborder or infraorder) Stylommatophora.
Terrestrial molluscs occur across most of the planet, with the exception of Antarctica and some islands. They inhabit a wide range of ecosystems, from deserts and tundras to rainforests.
In terms of survival, this group of species is currently one of the most threatened; there are more known species extinctions of terrestrial molluscs than in any other group of organisms. [4]
According to an estimate from Cameron, [4] of the 409 existing gastropod families, 119 include terrestrial molluscs. Among these 119 families, 104 are Stylommatophora, 7 are terrestrial pulmonates other than stylommatophorans, and 8 are operculates (formerly "prosobranchs", molluscs with an operculum, a group that primarily consists of marine snails).
"Prosobranchs"
"Pulmonates"
Athoracophoridae, common name the leaf-veined slugs, are a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the infraorder Stylommatophora, the stalk-eyed snails and slugs. Many of the species have an attractive pattern on their dorsal surface which resembles the veins in a leaf, hence the common name.
Agriolimacidae is a family of small and medium-sized land slugs, or shell-less snails, terrestrial pulmonate gastropod mollusks.
Zonitidae, common name the true glass snails, are a family of mostly rather small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Zonitoidea.
Limacidae, also known by their common name the keelback slugs, are a taxonomic family of medium-sized to very large, air-breathing land slugs, terrestrial pulmonate gastropod molluscs in the superfamily Limacoidea.
Endodontidae is a taxonomic family of very small air-breathing land snails and slugs, terrestrial pulmonate gastropod mollusks in the superfamily Punctoidea.
Helicarionidae is a family of air-breathing land snails or semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.
Euconulidae is a taxonomic family of minute, air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the superfamily Trochomorphoidea.
Subulininae is a subfamily of small tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Achatinidae.
Philomycidae are a family of air-breathing land slugs. They are terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea.
Monachoides incarnatus is a species of air-breathing land snail in the terrestrial pulmonate gastropod mollusk family Hygromiidae, the hairy snails and their allies.
Milacidae is a family of air-breathing, keeled, land slugs. These are shell-less terrestrial gastropod mollusks in the superfamily Parmacelloidea.
A love dart is a sharp, calcareous or chitinous dart which some hermaphroditic land snails and slugs create. Love darts are both formed and stored internally in a dart sac. These darts are made in sexually mature animals only, and are used as part of the sequence of events during courtship, before actual mating takes place. Darts are quite large compared to the size of the animal: in the case of the semi-slug genus Parmarion, the length of a dart can be up to one fifth that of the semi-slug's foot.
Ariophantidae is a taxonomic family of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.
Trochomorphidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea. Since 2017, its classification has been revised and it now belongs to the superfamily Trochomorphoidea
Staffordiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea.
Acavidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the superfamily Acavoidea.
The sensory organs of gastropods include olfactory organs, eyes, statocysts and mechanoreceptors. Gastropods have no sense of hearing.
The reproductive system of gastropods varies greatly from one group to another within this very large and diverse taxonomic class of animals. Their reproductive strategies also vary greatly, see Mating of gastropods.
The caudal mucous pit, or caudal mucous horn, is an anatomical structure on the tail end of the foot of various land snails and slugs, terrestrial pulmonate gastropod mollusks.
Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs to major rivers. The great majority of freshwater gastropods have a shell, with very few exceptions. Some groups of snails that live in freshwater respire using gills, whereas other groups need to reach the surface to breathe air. In addition, some are amphibious and have both gills and a lung. Most feed on algae, but many are detritivores and some are filter feeders.