The lobopodians, members of the informal group Lobopodia, or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as Aysheaia and Hallucigenia.
Hallucigenia is a genus of Cambrian animal resembling worms, known from articulated fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed as an enigmatic animal upside down and back to front. Hallucigenia is later recognized as part of lobopodians, a grade of Paleozoic panarthropods where the velvet worms, water bears, and arthropods arose.
A sclerite is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates.
Microdictyon is an extinct armoured worm-like animal coated with net-like scleritic plates, known from the Early Cambrian Maotianshan shale of Yunnan China and other parts of the world. Microdictyon is part of the ill-defined taxon – Lobopodia – that includes several other odd worm-like animals that resembling worm with legs, such as Hallucigenia, Onychodictyon, Cardiodictyon, Luolishania, and Paucipodia. The isolated sclerites of Microdictyon are known from other Lower Cambrian deposits. Microdictyon sclerites appear to have moulted; one sclerite seems to have been preserved during ecdysis.
Wiwaxia is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being 2 millimetres (0.079 in) long.
Anomalocaris is an extinct genus of radiodont (anomalocaridid), an order of animals thought to be closely related to ancestral arthropods. The first fossils of Anomalocaris were discovered in the Ogygopsis Shale by Joseph Frederick Whiteaves, with more examples found by Charles Doolittle Walcott in the Burgess Shale. Originally several fossilized parts discovered separately were thought to be three separate creatures, a misapprehension corrected by Harry B. Whittington and Derek Briggs in a 1985 journal article. Anomalocaris is thought to be one of the earliest examples of an apex predator, though others have been found in older Cambrian lagerstätten deposits.
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.
The Coelosclerithophorans are a polyphyletic group of organisms bearing hollow sclerites made of aragonite, and with a supposedly distinctive microstructure.
Orthrozanclus is a genus of sea creatures from the Middle Cambrian Burgess shale. Animals in this genus were one to two centimeters long, with spikes protruding from their armored bodies. The placement of this genus into a specific family is not universally accepted.
Fuxianhuia is a genus of Lower Cambrian fossil arthropod known from the Chengjiang fauna in China. Its purportedly primitive features have led to its playing a pivotal role in discussions about the euarthropod stem group. Nevertheless, despite being known from many specimens, disputes about its morphology, in particular its head appendages, have made it one of the most controversial of the Chengjiang taxa, and it has been discussed extensively in the context of the arthropod head problem.
Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade.
The Chancelloriids are an extinct family of animal common in sediments from the Early Cambrian to the early Late Cambrian. Many of these fossils consists only of spines and other fragments, and it is not certain that they belong to the same type of organism. Other specimens appear to be more complete and to represent sessile, bag-like organisms with a soft skin armored with star-shaped calcareous sclerites from which radiate sharp spines.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
Since 1990 there has been intense debate among paleontologists about the evolution in the Early Cambrian period of the "super-phylum" Lophotrochozoa, which is thought to include the modern molluscs, annelid worms and brachiopods, as well as their evolutionary "aunts" and "cousins".
Australohalkieria is an extinct genus of halkieriid from Australia and Antarctica.
Hurdia is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. As a radiodont like Peytoia and Anomalocaris, it is part of the ancestral lineage that led to euarthropods.
The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the late Silurian; they are mainly found as disarticulated sclerites, but are also preserved in many of the Cambrian lagerstätten. They take their name from the typifying genus Palaeoscolex. Other genera include Cricocosmia from the Lower Cambrian Chengjiang biota.
Sinosachites is a genus of 'halkieriid' known only from sclerites; these have internal chambers that are sub-perpendicular to the central canal, to which they are connected by narrow channels. The chambers are the same diameter, ~40 µm, as the longitudinal canals in Australohalkieria; their greater number and arrangement as lateral rather than longitudinal bodies reflects the greater size of the Sinosachites sclerites, which measure about 1–2 mm in length.
The camenellans, consisting of the genera Camenalla, Dailyatia, Kennardia, Kelanella and Lapworthella, are a group of Tommotiid invertebrates from the Cambrian period, reconstructed as sister to all others. They are known from isolated sclerites, but are believed to have a scleritomous, Halkieria-like construction.
Titanokorys is a genus of extinct hurdiid radiodont that existed during the middle part of the Cambrian. It is the largest member of its family from the Cambrian, with a body length of 50 centimeters long, making it one of the largest animals of the time. It bears a resemblance to a related genus Cambroraster. Based on the shape of its appendages, Titanokorys is speculated to have used them to sift through the sand looking for prey. Fossils of T. gainesi were first found within the Marble canyon in 2018. The fossils were not named until 2021 because they were assumed to be giant specimens of Cambroraster.
