Tornoceratidae

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Tornoceratidae
Temporal range: middle and upper Devonian
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Ammonoidea
Order: Goniatitida
Superfamily: Tornoceratoidea
Family: Tornoceratidae
Arthaber, 1911
Subfamilies

Tornoceratidae is a family of goniatitid ammonoids from the middle and upper Devonian. The family is included in the suborder Tornoceratina and the superfamily Tornoceratoidea.

Contents

Tornoceratids are subdiscoidal goniatitids with biconvex growth lines and sutures that form 6 to 10 lobes, the ventral one undivided, the lateral ones originating as subdivisions of external and internal lateral saddle. They are derived from the Anarcestida.

Taxonomy

Tornoceratidae as now understood is essentially the Tornoceratidae of Arthaber (1911) described in the American Treatise on Invertebrate Paleontology, Part L, revised to include three subfamilies, the Tornoceratinae, Aulatornoceratinae, and Falcitornoceratinae.

Miller et al. (1957) included the Tornoceratidae without subfamilies in the Cheilocerataceae, a superfaminily in the Goniatitida which also includes the Cheiloceratidae.

Tornoceratinae is based on the genus Tornoceras described by Alpheus Hyatt in 1884. Aulatornoceratinae and Falcitornoceratinae were established by Becker in 1993; Aulatornoceratinae based on Aulatornoceras , named by Schindewolf in 1922; Falcitornoceratinae on Falcitornoceras , named by House and Price in 1985.

The Tornoceratinae form the root stock of the tornoceratids and are derived from Anarcestida. Tornoceratinae in turn gave rise to Aulatornoceratinae and Falcitornoceratinae.

The Tornoceratinae includes Tornoceras, Epitornoceras, Lobotornoceras, Protornoceras, and Tornia, described in the Treatise (L 1957), along with Crassotornoceras, Linguatornoceras, Domanikoceras, Simcheiloceras, and Oxytornoceras which were added later.

Aulatornoceras, type for the Aulatornoceratinae was previously considered a subgenus of Tornoceras and is known from the Upper Devonian of North America and Europe. Aulatornoceras gave rise to Truyolsoceras and Polonoceras, which gave rise to Armatites, Kirsoceras, and Pernoceras, which gave rise to Pseudoclymenia making up the Aulotornoceratidae.

Genera making up the Falcitornoceratinae form an evolutionary series, Phoenixites, Falcitornoceras, Gundolficeras, Exotornoceras, leading up to Posttornoceras, type genus of the Posttornoceratidae.

Descendants

The Tornoceratidae probably gave rise to the Cheiloceratidae (Miller et al. 1964, fig 12, L27) but though which genus is unspecified (Saunders et al. 2004). The Tornoceratidae may also have given rise, though Tornoceras to the Clymeniida.

Related Research Articles

<span class="mw-page-title-main">Goniatite</span> Extinct order of molluscs

Goniatids, informally goniatites, are ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago. Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become extinct at the end of the Permian some 139 million years later.

Agathiceras is a subglobose goniatitid from the family Agathiceratidae, widespread and locally abundant in Lower Pennsylvanian to Middle Permian sediments, e.g. the Urals, Sicily, and Texas.

Aulatornoceras is a genus belonging to the subfamily Aulatornoceratinae, a member of the Goniatitida, an extinct order of shelled cephalopods included in the Ammonoidea. Aulatornoceras, which has been considered a subgenus of Tornoceras has ventro-lateral grooves. As with Tornoceras, the suture forms six lobes. The shell itself is involute.

<i>Armatites</i> Extinct genus of molluscs

Armatites is an Upper Devonian goniatitid included in the tornoceratid family. The shell, or conch, is discoidal, with flattened flanks, a flattened venter with a double keel, and a deep ventral sinus, but without ventrolateral grooves. The lateral saddle of the suture is broad. Armatites is known from the Upper Devonian (Fammenian) of Canning Basin in Australia. The subfamily in which it is included, the Aulatornoceratinae is also known from France with the genus Aulatornoceras.

Dimeroceratidae is one of three families in the Dimeroceratoidea, a goniatid superfamily included in the Ammonoidea; extinct shelled cephalopods with adorally convex septa and usually narrow ventro-marginal siphuncles.

Prolobitidae is a family of middle and upper Devonian ammonoid cephalopods currently included in the goniatitid suborder Tornoceratina and superfamily Dimeroceratoidea, but previously included in the ancestral Anarcestida.

Cheiloceratidae is a family of ammonoid cephalopods included in the goniatitid suborder Tornoceratina in which the suture has 4 to 12 lobes, the ventral one undivided and those in the lateral areas originating as subdivisions of internal and external lateral saddles.

Tornoceratoidea, also known as Tornocerataceae, is a superfamily of goniatitid ammonoids included in the suborder Tornoceratina. Tornoceratoidea, or Tornocerataceae, is essentially the Cheilocerataceae of Miller, Furnish, and Schindewolf (1957) in the Treatise Part L, revised to accommodate new taxa and new perspectives on the phylogeny.

The Posttornoceratidae are Late Devonian goniatites (Ammonoidea) included in the superfamily Tornoceratoidea. The family, Posttornoceratidae, named by Bogoslovsky in 1962, is based on the genus Posttornoceras, named by Wedekind in 1910, originally included in the Tornoceratidae.

Aulatornoceratinae is one of three subfamilies of the goniatitid family Tornoceratidae, an extinct order of Paleozoic ammonoid cephalopods. Aulotornoceratinae was established as a subfamily by R.T.Becker, 1993, initially for Aulatornoceras, named by Schindewolf, 1922. Subsequently, four other genera have been added.

Adrianitidae is a family in the Adrianitaceae, a superfamily of ammonites in the cephalopod order, Goniatitida, known from the Middle Pennsylvanian to the Middle Permian.

Adrianitinae is a subfamily of the Adrianitidae which is part of the goniatitid superfamily Adrianitaceae. The Adrianitinae which comprise the more advanced genera in the Adrianitidae have sutures that form 14 to 30 lobes. Shells may be discoidal or globular or in between.

<span class="mw-page-title-main">Prolecanitida</span> Extinct order of molluscs

Prolecanitida is an order of extinct ammonoid cephalopods, the major Late Paleozoic group of ammonoids alongside the order Goniatitida. Prolecanitids had narrow shells, discoidal (disc-shaped) to thinly lenticular (lens-shaped). They retained a retrochoanitic siphuncle, a simple form with septal necks extending backwards. As is typical for ammonoids, the siphuncle sits along the ventral margin of the shell.

<span class="mw-page-title-main">Agoniatitida</span> Extinct order of molluscs

Agoniatitida, also known as the Anarcestida, is the ancestral order within the cephalopod subclass Ammonoidea originating from bactritoid nautiloids, that lived in what would become Africa, Asia, Australia, Europe, and North America during the Devonian from about the lower boundary of Zlichovian stage into Taghanic event during upper middle Givetian, existing for approximately 25 million years.

<i>Cyclolobus</i>

Clyclolobus is a smooth, essentially involute subdiscoidal goniatitid ammonoid that has sutures with a bifurcate ventral lobe, flared outwardly at the end, in which the halves may be secondarily trifurcated, ending in sharp, narrow projections. Lateral sutural elements follow an acuate line that swings first to the front, then sharply to the rear before becoming hidden by the next whorl. Saddles are narrow, cumulous in appearance with short, irregular, rounded sub-endings. Ventro-lateal lobes are trifurcate with pointed, thorn-like projections.

Cravevoceras is an Upper Paleozoic ammonite in the goniatite family Cravenoceratidae, probably derived from Pachylyroceras and contemporary with other cravenoceratid genera like Caenolyroceras, Tympanoceras and later Alaoceras and Lyrogoniatites. It is also a member of the Neoglyphioceratoidea.

<span class="mw-page-title-main">Medlicottiinae</span> Extinct subfamily of molluscs

The Medlicottiinae is a subfamily of the Medlicottiidae, a family of ammonoid cephalopods included in the Prolecanitida, characterized by having discoidal to thinly lenticular shells with a retuse (grooved) venter and sutures with bifid auxiliary lobes.

<i>Tornoceras</i> Extinct genus of molluscs

Tornoceras is a strongly involute, subdiscoidal Middle and Upper Devonian goniatite with a suture that forms six to ten lobes. Tornoceras is an extinct genus of cephalopods that shares similarities with the modern pearly nautilus. This genus first appeared during the Devonian Period, which spanned from 416 million to 359 million years ago. The shell is circular and relatively flat, with the final whorl enveloping the previous ones. The sutures between the successive chambers of the shell exhibit a gently rippled pattern.

Phoenixites is an early genus of the Falcitornoceratinae, a subfamily of the goniatitid Tornoceratidae family. This genus was named by Becker in 1995. The type species is "Tornoceras" frechi.

Neoglaphyrites is a goniatitid ammonite that lived during the latest Pennsylvanian and early Permian. Its shell is ellipsoidal and moderately involute; the umbilicus deep and typically less than 15 per cent of the shell diameter but in some species closer to 20 per cent. Delicate growth lines forming ventral and lateral sinuses and ventrolateral and dorsolateral salients have been found on Canadian Arctic specimens. The suture is characterized by the ventral lobe split into two broad prongs that are separated by a high median ventral saddle; prongs closely approximate the width of the first lateral lobe. The first lateral saddle is evenly rounded and is nearly symmetrical. The umbilical lobe is V-shaped and internal lobes are deep and narrow.

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