The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.
The Vespidae are a large, diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps and many solitary wasps. Each social wasp colony includes a queen and a number of female workers with varying degrees of sterility relative to the queen. In temperate social species, colonies usually last only one year, dying at the onset of winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queens hibernate over winter in cracks or other sheltered locations. The nests of most species are constructed out of mud, but polistines and vespines use plant fibers, chewed to form a sort of paper. Many species are pollen vectors contributing to the pollination of several plants, being potential or even effective pollinators, while others are notable predators of pest insect species.
Wasps in the family Pompilidae are commonly called spider wasps, spider-hunting wasps, or pompilid wasps. The family is cosmopolitan, with some 5,000 species in six subfamilies. Nearly all species are solitary, and most capture and paralyze prey, though members of the subfamily Ceropalinae are kleptoparasites of other pompilids, or ectoparasitoids of living spiders.
The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps.
The Crabronidae are a large paraphyletic group of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. Several of the subfamilies of Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies.
The organ pipe mud dauber is a predatory wasp in the family Crabronidae. They are fairly large wasps, ranging from 3.9–5.1 cm, and have been recorded to fly from May to September. Female and male are similar in colour, a shiny black with pale yellow to white hind hindtarsomere. The organ pipe mud dauber feeds mainly on three genera of spider: Neoscona, Araneus, and Eustala. Melittobia, a parasitoid wasp, is a common ectoparasite of T. politum prepupae. Other sources of parasitism include the Bombyliid fly Anthrax, Chrysidid wasps, and various species of scavenger flies (Miltogramminae). The tufted titmouse is a known predator of T. politum, and may feed on them more commonly than previously thought as the holes made by the titmouse are similar in shape and size to those made by T. politum leaving the nest after pupation.
The Bembicinae comprise a large subfamily of crabronid wasps that includes over 80 genera and over 1800 species which have a worldwide distribution. They excavate nests in the soil, frequently in sandy soils, and store insects of several orders, for example Diptera, Orthoptera, Hemiptera, Lepidoptera and Odonata in the burrows. Some species are kleptoparasites of other Bembicinae. The different subgroups of Bembicinae are each quite distinctive, and rather well-defined, with clear morphological and behavioral differences between them.
Dryinidae is a cosmopolitan family of solitary wasps. Its name comes from the Greek drys for oak: Latreille named the type genus Dryinus because the first species was collected in an oak plant in Spain. The larvae are parasitoids of the nymphs and adults of Auchenorrhyncha. Dryinidae comprises approximately 1900 described species, distributed in 17 subfamilies and 53 genera.
The Stenogastrinae are a subfamily of social wasps included in the family Vespidae. They are sometimes called hover wasps owing to the particular hovering flight of some species. Their morphology and biology present interesting peculiarities.
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as their common ancestor is shared by bees and ants. Many wasps, those in the clade Aculeata, can sting their insect prey.
Ampulex is a large cosmopolitan genus of wasps belonging to the family Ampulicidae. Most of the >130 species occur in the tropics, particularly in the Old World, and fewer than 15 are known from the New World; fewer than 5 species are native to Europe or the United States, though the Old World species Ampulex compressa has spread to virtually everywhere that its host roaches can be found. The few species whose biology is known are parasitoids of cockroaches; they typically inject venom into the roach that subdues or immobilizes it, and then lay one to two eggs between the legs of the defenseless roach.
The thin-waisted social wasps, which typically reside in Neotropical regions from Central to South America, are a small genus of wasps that build nests and live in colonies ranging in size from 1 to 18 members. Microstigmus is widely considered to be the only true eusocial species within the family Crabronidae. Like all Hymenoptera, Microstigmus has an interesting sex determination pattern. Females are 2n (diploid), spawning from eggs that have been fertilized, while males are 1n (haploid) and spawn from unfertilized eggs. Female wasps contain the eggs within their egg sac and have the option of fertilizing them or not, thus having complete control of the gender of their offspring. This wasp genus, part of the suborder Apocrita, is distinguished physically by the narrow waist (petiole) between the end segment of the thorax (mesosoma) and the beginning of the abdomen (metasoma). Specifically, it is in the subgroup of "apoid wasps", those that exhibit social behavior in nesting and foraging. Evidence does exist that in some species social altruism is visible; however, studies on these topics have been limited. The small size and enclosed nest structure makes it difficult to obtain significant data. Although it was originally classified under the Sphecidae family due to its elongated petiole, Microstigmus has been reclassified into the new Crabronidae family.
Polistes carnifex is a neotropical vespid wasp in the cosmopolitan genus Polistes, known for its extremely painful and potent sting. The wasp is native to Central and South America. It is a very large yellow and brown paper wasp that establishes small colonies which build nests under the eaves of buildings or suspended from branches. The colonies are founded by solitary queens. Not all nests have a female with developed ovaries. Foraging adults bring nectar and macerated prey back to the nest to feed to the developing larvae which are individually housed in separate cells in the nest. It has mandibles with teeth.
Synoeca cyanea, commonly known as the marimbondo-tatu in Brazil, is a swarm-founding eusocial wasp. Native to Brazil and Argentina, S. cyanea is one of the largest and most aggressive species of social wasps and is feared in many rural areas. It begins its colony cycle in the early spring and continues until nest abandonment. Throughout its life, S. cyanea forage sugary substances and animal carcasses for food and wood pulp for its nest. S. cyanea is also known for its strong venom, which is enough to cause haemolytic activity.
Trypoxylon is a genus of wasps in the family Crabronidae. All Trypoxylon species that have been studied so far are active hunters of spiders, which they paralyse with a venomous sting, to provide as food to their developing larvae. Depending on the species, they will either construct their own nest from mud or find cavities that already exist. These cavities can range from keyholes to nail holes to previously abandoned nests, and are generally sealed with mud to create cells for their larvae.
Pison spinolae, commonly known as mason wasp, is a solitary wasp of the family Crabronidae, found throughout New Zealand.
Acrotaphus is a genus of ichneumon wasps in the family Ichneumonidae. There are about 26 described species in Acrotaphus.
Polychrus acutirostris, the Brazilian bush anole, is a species of lizard native to southern and eastern Brazil, Paraguay, Argentina, and eastern Bolivia. It is diurnal.
