Crabronidae | |
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Tachysphex sp. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Superfamily: | Apoidea |
Family: | Crabronidae Latreille, 1802 |
Subfamilies | |
Astatinae Contents |
The Crabronidae are a large paraphyletic group (nominally a family) of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of the Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. Several of the subfamilies of the Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies (example below).
This phylogenetic tree is based on Sann et al., 2018, which used phylogenomics to demonstrate that both the bees (Anthophila) and the Sphecidae arose from within the former Crabronidae, which is therefore paraphyletic, and which they suggested should be split into several families; the former family Heterogynaidae nests within the Bembicidae, as here defined. [1] These findings differ in several details from studies published by two other sets of authors in 2017, though all three studies demonstrate a paraphyletic "Crabronidae." [2] [3] Only three of these lineages were not included within Crabronidae in the past: Ampulicidae, Sphecidae, and Anthophila.
The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.
The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps.
The Spheciformes is a paraphyletic assemblage of insect families which collectively comprise the "sphecoid wasps". Larvae are carnivorous.
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic.
Cerceris is a genus of wasps in the family Philanthidae. It is the largest genus in the family, with 876 described species and 169 subspecies. The genus has a cosmopolitan distribution, with species on every continent.
Philanthidae is one of the largest families of wasp in the superfamily Apoidea, with 1167 species in 8 genera, most of which are Cerceris.
Heterogynaidae is a minor and disputed lineage of small spheciform wasps occurring in Madagascar, Botswana, Turkmenistan, Oman, the United Arab Emirates, and the Eastern Mediterranean area. The majority are dark in color and range in size from approximately 1.5 to 5.0 mm. Most specimens have been collected in arid climates, but one species from Madagascar is known to occur in a humid forest habitat. Although males have functional wings, heterogynaid females are typically brachypterous, a trait which is unique among spheciform wasps. Wing venation is reduced in both sexes. All species are diurnal, with the exception of H. nocticola. Other aspects of their biology are completely unknown, but details of their morphology have prompted researchers to hypothesize that they may be non-fossorial parasitoids adapted to hunt in tight spaces, such as under tree bark. This is speculative and has not yet been confirmed by actual observations of behavior. It is also possible that modifications of the female metasomal tergum VI and gonostyli may represent a unique prey transport mechanism, but this is also unconfirmed.
Mellinidae is a small family of wasps, comprising 17 described species in two genera. This group has traditionally been treated as a subfamily within Crabronidae (Mellininae), but recent phylogenomic studies have shown it to be a distinct family.
Pemphredonidae is a family of aphid wasps formerly treated as the subfamily Pemphredoninae. There are 19 genera and 556 described species in the family.
Astata is a cosmopolitan genus of solitary predatory wasps in the family Astatidae. They are known to prey on adults and nymphs of Pentatomidae. Astata is the largest genus in this subfamily, and is identified by features of its wing venation. The males of this genus and the related genus Dryudella have very large compound eyes that broadly meet at the top of the head.
Cerceris tuberculata is a species of wasp in the family Philanthidae.
Mimumesa is a genus of wasps in the family Psenidae. The species are found in the Holarctic. 32 species are known to exist.
Psenulus is a genus of wasps in the family Psenidae. The 173 species are found worldwide, but are best represented in the Indomalayan realm with 68. The Palearctic has 26, the Nearctic 4, and the Australasian realm 3. Psenulus is largely absent from South America and entirely absent from Melanesia and Polynesia. A recent phylogenetic analysis provided strong evidence that this genus is the closest living relative to bees.
Psenidae is a family of aphid wasps in the superfamily Apoidea formerly treated as the tribe Psenini. There are 12 genera and at least 485 described species of Psenidae.
Astata unicolor is a species of wasp in the family Astatidae. It is found in Central America and North America.
Astata bicolor is a species of wasp in the family Astatidae. It is found in Central America and North America.
Ammoplanops is a genus of aphid wasps in the family Ammoplanidae. There are about 15 species described in the genus Ammoplanops.
Pseneo punctatus is a species of aphid wasp in the family Psenidae. It is found in Central America and North America.
Dryudella caerulea is a species of wasp in the family Astatidae. It is found in Central America and North America.
Diploplectron ferrugineum is a species of wasp in the family Astatidae. It is found in North America.