Vespidae Temporal range: | |
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Vespula germanica | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Superfamily: | Vespoidea |
Family: | Vespidae Latreille, 1802 |
Subfamilies | |
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The Vespidae are a large (nearly 5000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps (such as Polistes fuscatus , Vespa orientalis , and Vespula germanica ) and many solitary wasps. [1] Each social wasp colony includes a queen and a number of female workers with varying degrees of sterility relative to the queen. In temperate social species, colonies usually last only one year, dying at the onset of winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queens hibernate over winter in cracks or other sheltered locations. The nests of most species are constructed out of mud, but polistines and vespines use plant fibers, chewed to form a sort of paper (also true of some stenogastrines). Many species are pollen vectors contributing to the pollination of several plants, being potential or even effective pollinators, [2] while others are notable predators of pest insect species, and a few species are invasive pests. [3] [4]
The subfamilies Polistinae and Vespinae are composed solely of eusocial species, while the Eumeninae, Euparagiinae, Gayellinae, Masarinae and Zethinae are all solitary with the exception of a few communal and several subsocial species. The Stenogastrinae are facultatively eusocial, considering nests may have one or several adult females; in cases where the nest is shared by multiple females (typically, a mother and her daughters) there is reproductive division of labor and cooperative brood care. [5]
In the Polistinae and Vespinae, rather than consuming prey directly, prey are premasticated and fed to the larvae, which in return, produce a clear liquid (with high amino acid content) for the adults to consume; the exact amino acid composition varies considerably among species, but it is considered to contribute substantially to adult nutrition. [6]
Fossils are known since Aptian of the Early Cretaceous, with several described species from Cretaceous amber. [7]
Hornets are the largest of the eusocial wasps, and are similar in appearance to yellowjackets, their close relatives. Some species can reach up to 5.5 cm (2.2 in) in length. They are distinguished from other vespine wasps by the relatively large top margin of the head. Worldwide, 22 species of Vespa are recognized. Most species only occur in the tropics of Asia, though the European hornet is widely distributed throughout Europe, Russia, North America, and north-eastern Asia. Wasps native to North America in the genus Dolichovespula are commonly referred to as hornets, but all of them are actually yellowjackets.
Paper wasps are a type of vespid wasps. The term is typically used to refer to members of the vespid subfamily Polistinae, though it often colloquially includes members of the subfamilies Vespinae and Stenogastrinae, which also make nests out of paper.
The European hornet is the largest eusocial wasp native to Europe. It is also the only true hornet found in North America, having been introduced to the United States and Canada from Europe as early as 1840. Vespines, such as V. crabro, are known for making intricate paper-like nests out of surrounding plant materials and other fibers. Unlike most other vespines, reproductive suppression involves worker policing instead of queen pheromone control, as was previously thought.
The subfamily Vespinae contains the largest and best-known groups of eusocial wasps, including true hornets, and the "yellowjackets". The remaining genus, Provespa, is a small, poorly known group of nocturnal wasps from Southeast Asia. One genus, Palaeovespa, has been described the Paleocene to Eocene fossil records of North America and Europe. Collectively, the group can be found on all continents except Antarctica, and several of these wasps are invasive species, introduced beyond their native ranges, and can be major pests.
Vespula vulgaris, known as the common wasp, is a species found in regions that include the United Kingdom, Ireland, Germany, India, China, New Zealand and Australia. It is sometimes known in English as the European wasp, but the same name is used for the species Vespula germanica or German wasp. In 2010, the ostensible Vespula vulgaris wasps in North America were found to be a different species, Vespula alascensis.
The Polistinae is a subfamily of eusocial wasps belonging to the family Vespidae. They are closely related to the wasps and true hornets of the subfamily Vespinae, containing four tribes. With about 1,100 species total, it is the second-most diverse subfamily within the Vespidae, and while most species are tropical or subtropical, they include some of the most frequently encountered large wasps in temperate regions.
The Stenogastrinae are a subfamily of social wasps included in the family Vespidae. They are sometimes called hover wasps owing to the particular hovering flight of some species. Their morphology and biology present interesting peculiarities.
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.
Polistes humilis, known as the Australian paper wasp, is a species of wasp in the family Vespidae that is found throughout Australia and which has been introduced to northern New Zealand. These paper wasps can be identified by their long thin legs and banded yellow and black coloring. They have been known to re-utilize old nests. While the species does not exhibit morphological class differences, there are distinct behavioral differences between queens and workers. In addition, the species is eusocial and benefits from relatedness between individuals. They are known for delivering a painful sting, especially when their nest is disturbed, a behavior that has been developed as a nest defense mechanism. While wasps are often viewed negatively, they play an important pollination role for many plants.
Polistes chinensis is a polistine vespid wasp in the cosmopolitan genus Polistes, and is commonly known as the Asian, Chinese or Japanese paper wasp. It is found in East Asia, in particular China and Japan. The subspecies P. chinensis antennalis is an invasive species in New Zealand, having arrived in 1979.
Vespula austriaca is an obligate parasitic wasp, parasitizing the nests of other species in the genus Vespula in the Old World. Its common host species include V. rufa in Europe, Japan, and East Siberia.V. austriaca wasps pollinate orchids.
Vespula squamosa, or the southern yellowjacket, is a social wasp. This species can be identified by its distinctive black and yellow patterning and orange queen. This species is typically found in eastern North America, and its territory extends as far south as Central America. Within these territories, they create enormous, multiple-comb nests. The colonies may be either annual or perennial depending on the climate, and in many perennial nests, polygyny takes place. In addition, this species uses pheromones both as a sexual attractant and an alarm signal. This species feeds on insects and animal carcasses; it does not produce honey. V. squamosa, a social insect, has developed a parasitic relationship with the species V. vidua and V. maculifrons. Due to their painful, venomous stings, the species is considered a pest.
Polistes carolina is one of two species of red paper wasp found in the eastern United States and is noted for the finer ridges on its propodeum. It is a social wasp in the family Vespidae and subfamily Polistinae. The species is native to the United States from Texas to Florida, north to New York, and west to Nebraska. The wasp's common name is due to the reddish-brown color of its head and body. P. carolina prefer to build their nests in protected spaces.
Polistes bellicosus is a social paper wasp from the order Hymenoptera typically found within Texas, namely the Houston area. Like other paper wasps, Polistes bellicosus build nests by manipulating exposed fibers into paper to create cells. P. bellicosus often rebuild their nests at least once per colony season due to predation.
Polistes japonicus is a eusocial paper wasp found in Japan. It was first described by Henri Louis Frédéric de Saussure in 1858. It is closely related to Polistes formosanus. This species lives in small colonies with few workers and a foundress queen. Nests of these wasps are sometimes used as a traditional medicine in Korea, China, and Japan.
Protopolybia chartergoides, also known as Pseudochartergus chartergoides, is a species of wasp within the genus Protopolybia. It is a social wasp found in southern Central America and northern South America.
Vespa affinis, the lesser banded hornet, is a common hornet in tropical and subtropical Asia.
Polistes bahamensis is a large species of colourful paper wasp in the genus Polistes of the family Vespidae which occurs in the Bahamas, Florida and Louisiana. It is also said to occur in Georgia.
Van der Vecht's gland or Van der Vecht's organ is a gland which is located in an area of modified cuticle on the rearmost gastral sternite of female wasps. This gland secretes chemicals which are important in the determination and maintenance of the hierarchy of groups of eusocial wasps and are used in the defence of the nests in others. In the Asian giant hornet the Van der Vecht's gland is used to scent mark hives of honey bees to attract other members of their colony to cooperatively attack the hive; the only known case of the gland's use to scent mark a food source. In the cleptoparasitic paper wasp Polistes semenowi the female usurps the host foundress, usually Polistes dominula and uses an enlarged Van der Vecht's gland to produce large quantities of hydrocarbons and to control the host workers, and even sometimes the host foundress. The gland was discovered by, and named in honour of, the Dutch entomologist Jacobus van der Vecht.
Vespula structor is a species of wasp in the family Vespidae found in India, Nepal, Laos, Myanmar, China and Bhutan.