Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamily Apoidea. They are currently considered a clade, called Anthophila. There are over 20,000 known species of bees in seven recognized biological families. Some species – including honey bees, bumblebees, and stingless bees – live socially in colonies while most species (>90%) – including mason bees, carpenter bees, leafcutter bees, and sweat bees – are solitary.
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism —that is, they have a wormlike larval stage and an inactive pupal stage before they mature.
Hornets are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to 5.5 cm (2.2 in) in length. They are distinguished from other vespine wasps by the relatively large top margin of the head. Worldwide, 22 species of Vespa are recognized. Most species only occur in the tropics of Asia, though the European hornet is widely distributed throughout Europe, Russia, North America, and north-eastern Asia. Wasps native to North America in the genus Dolichovespula are commonly referred to as hornets, but all of them are actually yellowjackets.
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" (petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host or in a nest cell provisioned by their mothers.
Halictidae is the second-largest family of bees with nearly 4,500 species. They are commonly called sweat bees, as they are often attracted to perspiration. Halictid species are an extremely diverse group that can vary greatly in appearance. These bees occur all over the world and are found on every continent except Antarctica. Usually dark-colored and often metallic, halictids are found in various sizes, colors and patterns. Several species are all or partly green and a few are red, purple, or blue. A number of them have yellow markings, especially the males, which commonly have yellow faces, a pattern widespread among the various families of bees. The family is one of many with short tongues and is best distinguished by the arcuate basal vein found on the wing. Females in this family tend to be larger than the males.
Wasps in the family Pompilidae are commonly called spider wasps, spider-hunting wasps, or pompilid wasps. The family is cosmopolitan, with some 5,000 species in six subfamilies. Nearly all species are solitary, and most capture and paralyze prey, though members of the subfamily Ceropalinae are kleptoparasites of other pompilids, or ectoparasitoids of living spiders.
Mason bee is a name now commonly used for species of bees in the genus Osmia, of the family Megachilidae. Mason bees are named for their habit of using mud or other "masonry" products in constructing their nests, which are made in naturally occurring gaps such as between cracks in stones or other small dark cavities. When available, some species preferentially use hollow stems or holes in wood made by wood-boring insects.
The term cuckoo bee is used for a variety of different bee lineages which have evolved the kleptoparasitic behaviour of laying their eggs in the nests of other bees, reminiscent of the behavior of cuckoo birds. The name is perhaps best applied to the apid subfamily Nomadinae, but is commonly used in Europe to mean bumblebees Bombus subgenus Psithyrus. Females of cuckoo bees are easy to recognize in almost all cases, as they lack pollen-collecting structures and do not construct their own nests. They often have reduced body hair, abnormally thick and/or heavily sculptured exoskeleton, and saber-like mandibles, although this is not universally true; other less visible changes are also common.
The Colletidae are a family of bees, and are often referred to collectively as plasterer bees or polyester bees, due to the method of smoothing the walls of their nest cells with secretions applied with their mouthparts; these secretions dry into a cellophane-like lining. The five subfamilies, 54 genera, and over 2000 species are all evidently solitary, though many nest in aggregations. Two of the subfamilies, Euryglossinae and Hylaeinae, lack the external pollen-carrying apparatus that otherwise characterizes most bees, and instead carry the pollen in their crops. These groups, and most genera in this family, have liquid or semiliquid pollen masses on which the larvae develop.
The Oxaeinae are an exclusively American subfamily of the bee family Andrenidae, consisting of large (13–26 mm), fast-flying bees, often with large eyes. The four constituent genera, with a total of 19 described species, range from the United States to Argentina. Some resources still use the name Oxaeidae, and treat them as a family, but they were moved to subfamily status in 1995.
Andrena is a genus of bees in the family Andrenidae. With over 1,500 species, it is one of the largest genera of animals. It is a strongly monophyletic group that is difficult to split into more manageable divisions; currently, Andrena is organized into 104 subgenera. It is nearly worldwide in distribution, with the notable exceptions of Oceania and South America. Bees in this genus are commonly known as mining bees due to their ground-nesting lifestyle.
The bee subfamily Andreninae is a nearly cosmopolitan lineage, with most of its diversity in one genus, Andrena, which contains over 1500 species. The remaining four genera in the subfamily only contain a total of 9 species.
Andrena agilissima is a species of mining bee. They are present in most of Europe, the Near East and North Africa and can be found from April through July. Andrena agilissima is an oligolectic species, feeding only on the pollen of a few genera of Cruciferous vegetables.
Vespula squamosa, or the southern yellowjacket, is a social wasp. This species can be identified by its distinctive black and yellow patterning and orange queen. This species is typically found in eastern North America, and its territory extends as far south as Central America. Within these territories, they create enormous, multiple-comb nests. The colonies may be either annual or perennial depending on the climate, and in many perennial nests, polygyny takes place. In addition, this species uses pheromones both as a sexual attractant and an alarm signal. This species feeds on insects and animal carcasses; it does not produce honey. V. squamosa, a social insect, has developed a parasitic relationship with the species V. vidua and V. maculifrons. Due to their painful, venomous stings, the species is considered a pest.
Andrena antoinei is an extinct species of mining bee in the family Andrenidae described from a single fossil found in a Late Oligocene lake in present-day France that existed in semi-arid conditions.
Andrena scotica, the chocolate mining bee or hawthorn bee, is a species of mining bee from the family Andrenidae. It occurs in western Europe and is one of the most frequently encountered mining bees found in Great Britain, where it had been previously misidentified as Andrena carantonica.
The Appalachian miner bee is a species of miner bee in the family Andrenidae. Another common name for this species is Alleghany andrena. It is found in North America. Its nests typically only have one or two cells. Adults overwinter in soil.
Andrena obscuripennis is a species of mining bee in the family Andrenidae. It is found in North America.
Andrena prunorum, otherwise known as the purple miner bee, is a species of solitary bees in the family Andrenidae. It is commonly found in the continental United States as well as much of North and Central America. Andrena prunorum is a spring-flying, ground-nesting bee that serves as a ubiquitous generalist in ecological settings. Both males and females live as prepupae in the winter in which they mate, and the females seek new sites for ground burrows. From there, they construct small cells surrounding a ball of pollen combined with nectar to nourish a laid egg before each cell is sealed, and the cycle begins anew. A. prunorum generally prefer the pollen derived from Rosaceae plants but will pollinate fruit trees if given the opportunity.
Megalopta is a widespread neotropical genus of bees in the tribe Augochlorini in family Halictidae, known as the sweat bees. They are the largest of the five nocturnal genera in Augochlorini. Most have pale integumentary pigmentation, and all have large ocelli, most likely a feature of their nocturnal behavior. They live in tropical Central America and the entirety of South America. The subgenus Noctoraptor is cleptoparasitic. They are not known from the fossil record.
