The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" (petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host or in a nest cell provisioned by their mothers.
The superfamily Ichneumonoidea contains one extinct and three extant families, including the two largest families within Hymenoptera: Ichneumonidae and Braconidae. The group is thought to contain as many as 100,000 species, many of which have not yet been described. Like other parasitoid wasps, they were long placed in the "Parasitica", variously considered as an infraorder or an unranked clade, now known to be paraphyletic.
Vespoidea is a superfamily of wasps in the order Hymenoptera. Vespoidea includes wasps with a large variety of lifestyles including eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores.
The Crabronidae are a large paraphyletic group of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of the Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. Several of the subfamilies of the Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies.
The Spheciformes is a paraphyletic assemblage of insect families which collectively comprise the "sphecoid wasps". Larvae are carnivorous.
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders.
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic.
Proctotrupoidea is a hymenopteran superfamily containing seven extant families, though others have been recognized in the past, most of these having been removed to a recently erected superfamily Diaprioidea. Of the remaining families, only Proctotrupidae contains a substantial number of species, with over 400 described. The others are small, often relictual groups. See links for individual families for details of life history and diversity.
Heterogynaidae is a minor and disputed lineage of small spheciform wasps occurring in Madagascar, Botswana, Turkmenistan, Oman, the United Arab Emirates, and the Eastern Mediterranean area. The majority are dark in color and range in size from approximately 1.5 to 5.0 mm. Most specimens have been collected in arid climates, but one species from Madagascar is known to occur in a humid forest habitat. Although males have functional wings, heterogynaid females are typically brachypterous, a trait which is unique among spheciform wasps. Wing venation is reduced in both sexes. All species are diurnal, with the exception of H. nocticola. Other aspects of their biology are completely unknown, but details of their morphology have prompted researchers to hypothesize that they may be non-fossorial parasitoids adapted to hunt in tight spaces, such as under tree bark. This is speculative and has not yet been confirmed by actual observations of behavior. It is also possible that modifications of the female metasomal tergum VI and gonostyli may represent a unique prey transport mechanism, but this is also unconfirmed.
Mellinidae is a small family of wasps, comprising 17 described species in two genera. This group has traditionally been treated as a subfamily within Crabronidae (Mellininae), but recent phylogenomic studies have shown it to be a distinct family.
Bothynostethini is a small neotropical tribe of solitary wasps partly recognizable for having peculiar modifications at the apex of the hind femora.
The subfamily Pemphredoninae also known as the aphid wasps, is a large group in the wasp family Crabronidae, with over 1000 species. Historically, this subfamily has frequently been accorded family status. In some recent phylogenetic analyses, one of the subtribes within this group is the sister lineage to the superfamily Apoidea, and accorded family rank as Ammoplanidae along with Pemphredonidae and Psenidae so as to keep families monophyletic.
Ammophila sabulosa, the red-banded sand wasp, is a species of the subfamily Ammophilinae of the solitary hunting wasp family Sphecidae, also called digger wasps. Found across Eurasia, the parasitoid wasp is notable for the mass provisioning behaviour of the females, hunting caterpillars mainly on sunny days, paralysing them with a sting, and burying them in a burrow with a single egg. The species is also remarkable for the extent to which females parasitise their own species, either stealing prey from nests of other females to provision their own nests, or in brood parasitism, removing the other female's egg and laying one of her own instead.
Ammophilinae is a subfamily of thread-waisted wasps in the family Sphecidae. There are about 6 genera and more than 320 described species in Ammophilinae.
Psenulus is a genus of wasps in the family Crabronidae. The 160 species are found worldwide, but are best represented in the Indomalayan realm with 68. The Palearctic has 26, the Nearctic 4, and the Australasian realm 3. Psenulus is largely absent from South America and entirely absent from Melanesia and Polynesia. A recent phylogenetic analysis provided strong evidence that this genus is the closest living relative to bees.
The Chyphotidae are a family of wasps similar to the Mutillidae, differing most visibly in the presence, in females, of a suture separating the pronotum from the mesonotum. These species are found primarily in arid regions in the southwestern United States and adjacent regions in Mexico.
Diaprioidea is a hymenopteran superfamily containing five extant families, though in the past these families were included in the superfamily Proctotrupoidea.
Psenini is a tribe of aphid wasps in the family Crabronidae. There are about 11 genera and at least 460 described species in Psenini.
Ammoplanidae is a family of aphid wasps formerly treated as the Crabronidae subtribe Ammoplanina. There are about 10 genera and at least 130 described species in Ammoplanina. Phylogenetic analyses in 2018 and 2021 have confirmed this group as the sister lineage to the bees, and thus accorded the group family rank.
