Spheciformes | |
---|---|
![]() | |
Ampulex compressa | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Superfamily: | Apoidea |
(unranked): | Spheciformes |
Families [1] | |
The Spheciformes is a paraphyletic assemblage of insect families which collectively comprise the "sphecoid wasps", and includes about 10 000 species. Larvae are carnivorous.
Spheciformes includes all the members of the superfamily Apoidea which are not bees and which in older classifications were called the "Sphecoidea". Some are also described as mud daubers.
The group is paraphyletic because bees are believed to have arisen from a subgroup within the family Ammoplanidae, thus Spheciformes does not include all of the descendants of its common ancestor. [2]
Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamily Apoidea. They are currently considered a clade, called Anthophila. There are over 20,000 known species of bees in seven recognized biological families. Some species – including honey bees, bumblebees, and stingless bees – live socially in colonies while most species (>90%) – including mason bees, carpenter bees, leafcutter bees, and sweat bees – are solitary.
Apidae is the largest family within the superfamily Apoidea, containing at least 5700 species of bees. The family includes some of the most commonly seen bees, including bumblebees and honey bees, but also includes stingless bees, carpenter bees, orchid bees, cuckoo bees, and a number of other less widely known groups. Many are valuable pollinators in natural habitats and for agricultural crops.
The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" (petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host or in a nest cell provisioned by their mothers.
Vespoidea is a superfamily of wasps in the order Hymenoptera. Vespoidea includes wasps with a large variety of lifestyles including eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores.
The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps.
The Crabronidae is a large family of wasps within the superfamily Apoidea.
The subfamily Dasypodainae is a small subfamily of melittid bees, with more than 100 species in eight genera, found in Africa and the northern temperate zone, primarily in xeric habitats.
Meganomiinae is a subfamily of melittid bees, with 10 species in four genera, found only in Africa, primarily in xeric habitats, with the distributional limits in Yemen and Madagascar. They are rather different in appearance from the other groups of past/present melittids, being large bees (10–22 mm), mostly black with strong yellow markings, resembling anthidiine megachilids. Males of this subfamily are known to have hidden sterna.
Cerceris is a genus of wasps in the family Philanthidae. It is the largest genus in the family, with 876 described species and 169 subspecies. The genus has a cosmopolitan distribution, with species on every continent.
Philanthidae is one of the largest families of wasp in the superfamily Apoidea, with 1167 species in 8 genera, most of which are Cerceris.
Heterogynaidae is a minor and disputed lineage of small spheciform wasps occurring in Madagascar, Botswana, Turkmenistan, Oman, the United Arab Emirates, and the Eastern Mediterranean area. The majority are dark in color and range in size from approximately 1.5 to 5.0 mm. Most specimens have been collected in arid climates, but one species from Madagascar is known to occur in a humid forest habitat. Although males have functional wings, heterogynaid females are typically brachypterous, a trait which is unique among spheciform wasps. Wing venation is reduced in both sexes. All species are diurnal, with the exception of H. nocticola. Other aspects of their biology are completely unknown, but details of their morphology have prompted researchers to hypothesize that they may be non-fossorial parasitoids adapted to hunt in tight spaces, such as under tree bark. This is speculative and has not yet been confirmed by actual observations of behavior. It is also possible that modifications of the female metasomal tergum VI and gonostyli may represent a unique prey transport mechanism, but this is also unconfirmed.
Astatidae is a cosmopolitan family of solitary wasps, peculiar for their males having very large compound eyes that broadly meet at the top of the head. The largest genus in this family is Astata, with about half of more than 160 species in the family.
Mellinidae is a small family of wasps, comprising 17 described species in two genera. This group has traditionally been treated as a subfamily within Crabronidae (Mellininae), but recent phylogenomic studies have shown it to be a distinct family.
Pemphredonidae is a family of aphid wasps formerly treated as the subfamily Pemphredoninae. There are 19 genera and 556 described species in the family.
Melittidae is a small bee family, with over 200 described species in three subfamilies. The family has a limited distribution, with all described species restricted to Africa and the northern temperate zone.
Psenidae is a family of aphid wasps in the superfamily Apoidea formerly treated as the tribe Psenini. There are 12 genera and at least 485 described species of Psenidae.
Cerceris clypeata is a species of weevil wasp in the family Philanthidae. It is found in North America.
Mimesa is a genus of wasps belonging to the family Psenidae. The species of this genus are found in Europe and North America.
Bembicinae is one of the two subfamilies of Bembicidae, along with Nyssoninae.