| Alatochelon Temporal range: | |
|---|---|
 | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | Testudines |
| Suborder: | Cryptodira |
| Superfamily: | Testudinoidea |
| Family: | Testudinidae |
| Genus: | † Alatochelon Pérez-García, Vlachos, & Murelaga, 2020 |
| Species: | †A. myrteum |
| Binomial name | |
| †Alatochelon myrteum Pérez-García, Vlachos, & Murelaga, 2020 | |
Alatochelon is an extinct genus of tortoise with a single known species, Alatochelon myrteum. [1]
Alatochelon is derived from the Ancient Greek alato-, meaning salt; and -chelon, meaning turtle. The 'salt' is in reference to the Messinian salinity crisis, during which it is hypothesized that the African spurred tortoise may have crossed over into Europe, and differentiated into Alatochelon myrteum through allopatric speciation. [2] Myrteum is derived from the Latin word 'myrtle', which is believed to be the ancient name of Murica, the region of Spain in which the first fossil of Alatochelon myrteum was found. [2]
Alatochelon myrteum was a large species of tortoise that lived in the lower Pliocene era. [2] As of 2023, the sole fossil found came from the Region of Murcia, Spain; thus its exact range is unknown. Alatochelon myrteum had a tall carapace, and a maximum shell length of at least one meter. [2] Other defining characteristics include a longer-than-wide nuchal scute and a well developed nuchal notch. [2]
Alatochelon myrteum was likely herbivorous. [3]
The Pliocene is the epoch in the geologic time scale that extends from 5.333 million to 2.58 million years ago. It is the second and most recent epoch of the Neogene Period in the Cenozoic Era. The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian Stage, which lasted from 2.588 to 1.806 million years ago, and is now included in the Pleistocene.
Tortoises are reptiles of the family Testudinidae of the order Testudines. Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backward into the shell to protect them.
The Messinian salinity crisis was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (drying-up) throughout the latter part of the Messinian age of the Miocene epoch, from 5.96 to 5.33 Ma. It ended with the Zanclean flood, when the Atlantic reclaimed the basin.
In biogeography, the Mediterranean Basin, also known as the Mediterranean Region or sometimes Mediterranea, is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and warm to hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation. It was a very important part of Mediterranean civilizations.
The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma. It follows the Tortonian and is followed by the Zanclean, the first age of the Pliocene.
Nuralagus is an extinct genus of leporid, with a single species, Nuralagus rex, described in 2011. It lived on Menorca, one of the Balearic Islands in the western Mediterranean during the Pliocene epoch. It is the largest known lagomorph to have ever existed, with an estimated weight of 8–12 kilograms (18–26 lb), nearly double the weight of the average Flemish Giant rabbit. It likely went extinct at the Pliocene-Pleistocene transition when Mallorca and Menorca were united as one island, letting the mammalian fauna of Mallorca, including the goat-like ungulate Myotragus, colonize Nuralagus's habitat.
Stupendemys is an extinct genus of freshwater side-necked turtle, belonging to the family Podocnemididae. It is the largest freshwater turtle known to have existed, with a carapace over 2 meters long. Its fossils have been found in northern South America, in rocks dating from the Middle Miocene to the very start of the Pliocene, about 13 to 5 million years ago. Male specimens are known to have possessed bony horns growing from the front edges of the shell and the discovery of the fossil of a young adult shows that the carapace of these turtles flattens with age. A fossil skull described in 2021 indicates that Stupendemys was a generalist feeder.
Pliohyrax, is a genus of hyracoids. It grew to sizes greatly exceeding those of any living hyrax, though it was by no means the largest member of this family.
Asoriculus is an extinct genus of terrestrial shrews in the subfamily Soricinae and tribe Nectogalini, native to Europe and North Africa.
Prolagus is an extinct genus of lagomorph. Over 20 species have been named, and the genus was abundant and widespread in Europe during the Neogene. However, by the end of the Middle Pleistocene, it was confined to a single species, the Sardinian pika, on the Corsica, Sardinia, and surrounding islands, where it survived into historical times. In North Africa and Western Asia, the genus is known from the Miocene and Pliocene. The scientific name may mean "before hares" or "primitive hares". Its taxonomy is disputed, with it either being considered a member of the family Ochotonidae, which includes living pikas, or the only member of the family Prolagidae.
Metaxytherium is an extinct genus of dugong that lived from the Oligocene until the end of the Pliocene. Fossil remains have been found in Africa, Europe, North America and South America. Generally marine seagrass specialists, they inhabited the warm and shallow waters of the Paratethys, Mediterranean, Caribbean Sea and Pacific coastline. American species of Metaxytherium are considered to be ancestral to the North Pacific family Hydrodamalinae, which includes the giant Steller's sea cow.
Aldabrachampsus is an extinct genus of small horned crocodile known from fragmentary remains. It lived during the Pleistocene on Aldabra Atoll, Seychelles in the western Indian Ocean. The name Aldabrachampsus dilophus means "Two-crested crocodile from Aldabra". It was a small animal, reaching a length of 2–2.5 m, comparable in size to the smallest extant crocodilians.
A gazelle is one of many antelope species in the genus Gazella. There are also seven species included in two further genera; Eudorcas and Nanger, which were formerly considered subgenera of Gazella. A third former subgenus, Procapra, includes three living species of Asian gazelles.
Samu is the nickname given to a fragmentary Middle Pleistocene human occipital, also known as Vertesszolos Man or Vertesszolos occipital, discovered in Vértesszőlős, Central Transdanubia, Hungary.
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. Herman Melville often referred to whales as "Leviathans" in his book. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, the United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Pterodromoides is an extinct genus of fulmarine petrel dating from the Late Miocene. It contains a single species, P. minoricensis. Its fossil remains were first discovered at the Punta Nati palaeontological site on the island of Menorca in the Balearic archipelago of the western Mediterranean. An additional specimen from North Carolina, USA has also been referred to this species, suggesting it lived across the North Atlantic. It was described in 2001, with the authors justifying the creation of a new genus by the large orbitonasal opening and characters of the postcranial skeleton, despite the similarity of the cranial osteology to that of Pagodroma.
Crocodylus falconensis is an extinct species of crocodile known from the early Pliocene of the lower part of the Vergel Member of the San Gregorio Formation of Venezuela. C. falconensis was named in 2013 after Falcón State and is thought to be the basalmost species of Crocodylus found in the Neotropics.
Titanochelon is an extinct genus of giant tortoises known from the Early Miocene to the beginning of the Pleistocene in Europe, extending from the Iberian Peninsula to Anatolia. Some members of the genus were larger than extant giant tortoises, with a shell length of up to 2 m.
Osodobenus is an extinct genus of walrus from the Miocene to Pliocene of California. Osodobenus may have been the first tusked walrus and shows several adaptations that suggest it was a suction feeder, possibly even a benthic feeder like modern species. Three skulls are known showing pronounced sexual dimorphism, with the female lacking the same tusks as the male. Only a single species, Osodobenus eodon, is currently recognized.
Solitudo is an extinct genus of tortoise that was found during the Pliocene and Pleistocene on the Mediterranean islands of Menorca, Malta and Sicily. The genus includes three described species, Solitudo robusta, Solitudo gymnesica and Solitudo sicula as well as a likely fourth, undescribed species from Monte Pellegrino in Sicily. Solitudo sicula, the youngest of the species, died out approximately 12.5 thousand years BP. The largest species, Solitudo gymnesica, has been estimated to have reached a carapace length of 1.1–1.3 m (3.6–4.3 ft).