Anoplolepis custodiens | |
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Anoplolepis custodiens worker | |
Scientific classification ![]() | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | Formicinae |
Tribe: | Lasiini |
Genus: | Anoplolepis |
Species: | A. custodiens |
Binomial name | |
Anoplolepis custodiens | |
Anoplolepis custodiens, commonly known as the common pugnacious ant, [2] is a species of ant in the genus Anoplolepis , native to central and southern Africa. Individual ants may come in a range of colors and sizes. Found in dry and cultivated areas, it is an important source of prey for some pangolins and aardvarks. Although generally aggressive, ants from other colonies of Anoplolepis custodiens are not attacked.
The worker common pugnacious ant varies greatly in size and is up to 10 mm (0.4 in) long, and polymorphic, being yellowish-brown, reddish-brown or darker chocolate-brown, often with the abdomen darker than the rest of the ant. Darker individuals can be distinguished from the black pugnacious ant (Anoplolepis steingroeveri) by the chequer-board dark pattern on the gaster, which is caused by reflection of light by pubescent hairs which lie in different directions on the two sides. [2]
This ant is native to tropical sub-Saharan Africa. Its range includes Angola, Botswana, Democratic Republic of Congo, Eswatini, Ethiopia, Lesotho, Mozambique, Namibia, South Africa, Tanzania, Zambia, Zanzibar and Zimbabwe. It is a terrestrial species living in dry, open habitats such as the Karoo in South Africa, and is found in cultivated areas and vineyards. [2]
Anoplolepis custodiens is a very aggressive ant and tends to dominate ant communities wherever it is found; nevertheless, it has been shown to be tolerant towards ants of its own species from other colonies. [2] This fact may facilitate its invasive expansion locally. It is a generalist feeder on dead and living animal material, nectar and honeydew. [2] It has been found that both larger and smaller workers forage on the ground, whereas it is almost exclusively smaller workers that forage in bushes, indicating the possibility of workers adopting size-related roles. [3]
A study of pangolins in Sabi Sand Game Reserve in South Africa showed that they fed exclusively on ants and termites and that A. custodiens constituted 77% of their diet, mostly dug out from shallow subterranean locations. [4] Similarly, A. custodiens forms a high proportion of the diet of aardvarks, being the most prominent prey species, but this largely reflects the fact that it is the most abundant ant species in the area. [5]
The aardvark is a medium-sized, burrowing, nocturnal mammal native to Africa. It is the only living species of the order Tubulidentata, although other prehistoric species and genera of Tubulidentata are known. Unlike most other insectivores, it has a long snout, similar to that of a pig, which is used to sniff out food.
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists.
Termites are a group of detritophagous eusocial insects which consume a wide variety of decaying plant material, generally in the form of wood, leaf litter, and soil humus. They are easily identified by the typically soft-bodied and unpigmented worker caste for which they have been colloquially termed "white ants"; however they are not ants to which they are distantly related. About 2,972 extant species are currently described, 2,105 of which are members of the family Termitidae.
The ground pangolin, also known as Temminck's pangolin, Cape pangolin or steppe pangolin, is one of four species of pangolins which can be found in Africa, and the only one in southern and eastern Africa. The animal was named for the Dutch zoologist Coenraad Jacob Temminck. As a group, pangolins are among the most critically endangered and illegally trafficked animals in the world.
The bat-eared fox is a species of fox found on the African savanna. It is the only extant species of the genus Otocyon and considered a basal canid species. Fossil records indicate this canid first appeared during the middle Pleistocene.
Weaver ants or green ants are eusocial insects of the family Formicidae. Weaver ants live in trees and are known for their unique nest building behaviour where workers construct nests by weaving together leaves using larval silk. Colonies can be extremely large consisting of more than a hundred nests spanning numerous trees and containing more than half a million workers. Like many other ant species, weaver ants prey on small insects and supplement their diet with carbohydrate-rich honeydew excreted by small insects (Hemiptera). Weaver ant workers exhibit a clear bimodal size distribution, with almost no overlap between the size of the minor and major workers. The major workers are approximately 8–10 mm (0.31–0.39 in) in length and the minors approximately half the length of the majors. Major workers forage, defend, maintain, and expand the colony whereas minor workers tend to stay within the nests where they care for the brood and 'milk' scale insects in or close to the nests.
The Indian pangolin, also called thick-tailed pangolin and scaly anteater is a pangolin native to the Indian subcontinent. Like other pangolins, it has large, overlapping scales on its body which act as armour. The colour of its scales varies depending on the colour of the earth in its surroundings. It can also curl itself into a ball as self-defence against predators such as the tiger. It is an insectivore feeding on ants and termites, digging them out of mounds and logs using its long claws, which are as long as its fore limbs. It is nocturnal and rests in deep burrows during the day.
The name army ant (or legionary ant or marabunta) is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limited area.
The banded mongoose is a mongoose species native from the Sahel to Southern Africa. It lives in savannas, open forests and grasslands and feeds primarily on beetles and millipedes. Mongooses use various types of dens for shelter including termite mounds. While most mongoose species live solitary lives, the banded mongoose live in colonies with a complex social structure.
Eciton burchellii is a species of New World army ant in the genus Eciton. This species performs expansive, organized swarm raids that give it the informal name, Eciton army ant. This species displays a high degree of worker polymorphism. Sterile workers are of four discrete size-castes: minors, medias, porters (sub-majors), and soldiers (majors). Soldiers have much larger heads and specialized mandibles for defense. In lieu of underground excavated nests, colonies of E. burchellii form temporary living nests known as bivouacs, which are composed of hanging live worker bodies and which can be disassembled and relocated during colony emigrations. Eciton burchellii colonies cycle between stationary phases and nomadic phases when the colony emigrates nightly. These alternating phases of emigration frequency are governed by coinciding brood developmental stages. Group foraging efforts known as "raids" are maintained by the use of pheromones, can be 200 metres (660 ft) long, and employ up to 200,000 ants. Workers are also adept at making living structures out of their own bodies to improve efficiency of moving as a group across the forest floor while foraging or emigrating. Workers can fill "potholes" in the foraging trail with their own bodies, and can also form living bridges. Numerous antbirds prey on the Eciton burchellii by using their raids as a source of food. In terms of geographical distribution, this species is found in the Amazon jungle and Central America.
Iridomyrmex is a genus of ants called rainbow ants first described by Austrian entomologist Gustav Mayr in 1862. He placed the genus in the subfamily Dolichoderinae of the family Formicidae. It has 79 described species and five fossil species. Most of these ants are native to Australia; others are found in Asia and Oceania, and they have been introduced to Brazil, New Zealand, and the United Arab Emirates. Fossil species are known from China, France, and the United States.
The meat ant, also known as the gravel ant or southern meat ant, is a species of ant endemic to Australia. A member of the genus Iridomyrmex in the subfamily Dolichoderinae, it was described by British entomologist Frederick Smith in 1858. The meat ant is associated with many common names due to its appearance, nest-building behaviour and abundance, of which its specific name, purpureus, refers to its coloured appearance. It is among the best-known species of ant found throughout Australia; it occurs in almost all states and territories except for Tasmania. Its enormous distribution, aggression and ecological importance have made this ant a dominant species.
Myrmecophagy is a feeding behavior defined by the consumption of termites or ants, particularly as pertaining to those animal species whose diets are largely or exclusively composed of said insect types. Literally, myrmecophagy means "ant eating" rather than "termite eating". The two habits often overlap, as both of these eusocial insect types often live in large, densely populated nests requiring similar adaptations in the animal species that exploit them.
Myrmecia nigriceps, also known as the black-headed bull ant, is a species of ant endemic to Australia. A member of the genus Myrmecia in the subfamily Myrmeciinae, it was first described by Austrian entomologist Gustav Mayr in 1862. These ants are large, varying from 19 to 23 millimetres in length. However, colonies contain workers that are much smaller, usually half the size of normal workers. The queens are the largest while the males are the smallest, which can be easily identified due to their small mandibles.
The green-head ant, also known as the green ant or metallic pony ant, is a species of ant that is endemic to Australia. It was described by British entomologist Frederick Smith in 1858 as a member of the genus Rhytidoponera in the subfamily Ectatomminae. These ants measure between 5 to 7 mm. The queens and workers look similar, differing only in size, with the males being the smallest. They are well known for their distinctive metallic appearance, which varies from green to purple or even reddish-violet. Among the most widespread of all insects in Australia, green-head ants are found in almost every Australian state, but are absent in Tasmania. They have also been introduced in New Zealand, where several populations have been established.
Nasutitermes corniger is a species of arboreal termite that is endemic to the neotropics. It is very closely related to Nasutitermes ephratae. The species has been studied relatively intensively, particularly on Barro Colorado Island, Panama. These studies and others have shown that the termite interacts with many different organisms including a bat that roosts in its nest and various species of ants that cohabit with the termite.
The Hodotermitidae are a basal Old World family of termites known as the Harvester termites. They are distinguished by the serrated inner edge of their mandibles, and their functional compound eyes which are present in all castes. They forage for grass at night and during daylight hours, and the pigmented workers are often observed outside the nest. Their range includes the deserts and savannas of Africa, the Middle East, and Southwest Asia. Their English name refers to their habit of collecting grass, which is not unique to the family however.
Trichomyrmex destructor is a species of ant in the subfamily Myrmicinae. Its common names include destructive trailing ant or Singapore ant. It is a pest species in urban areas, known for causing costly damage to structures, vehicles, and electronic devices with its chewing activity. In 2015, the species was moved from the genus Monomorium to the revised genus Trichomyrmex.
Anoplolepis steingroeveri is a species of ant. It is commonly known as the black pugnacious ant. It is native to southern Africa, and occurs in South Africa, Lesotho, Botswana and Namibia. The worker is similar in appearance to darker morphs of the common pugnacious ant, but that species has a chequer-board-like dark pattern on the gaster, which is caused by reflection of light by pubescent hairs which lie in different directions on the two sides.
An ant supercolony is an exceptionally large ant colony, consisting of a high number of spatially separated but socially connected nests of a single ant species, spread over a large area without territorial borders. Supercolonies are typically polygynous, containing many egg-laying females. Workers and queens from different nests within the same supercolony can freely move among the nests, and all workers cooperate indiscriminately with each other in collecting food and care of the brood, and show no apparent mutual aggressive behavior.