In mammals, invertebrates and most fish, [1] [2] the anus (pl.: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the end of the rectum, i.e. the opposite end from the mouth. Its function is to facilitate the expulsion of wastes that remain after digestion.
Bowel contents that pass through the anus include the gaseous flatus and the semi-solid feces, which (depending on the type of animal) include: indigestible matter such as bones, hair pellets, endozoochorous seeds and digestive rocks; [3] residual food material after the digestible nutrients have been extracted, for example cellulose or lignin; ingested matter which would be toxic if it remained in the digestive tract; excreted metabolites like bilirubin-containing bile; and dead mucosal epithelia or excess gut bacteria and other endosymbionts. Passage of feces through the anus is typically controlled by muscular sphincters, and failure to stop unwanted passages results in fecal incontinence.
Amphibians, reptiles and birds use a similar orifice (known as the cloaca) for excretion, copulation, and egg-laying. Monotreme mammals also have a cloaca, which is thought to be a feature inherited from the earliest amniotes. Marsupials have a single orifice for excreting both solids and liquids and, in females, a separate vagina for reproduction. Female placentals have completely separate orifices for defecation, urination, and reproduction; males have one opening for defecation and another for both urination and reproduction, although the channels flowing to that orifice are almost completely separate.
The development of the anus was an important stage in the evolution of multicellular animals. It appears to have happened at least twice, following different paths in protostomes and deuterostomes. This accompanied or facilitated other important evolutionary developments: the bilaterian body plan, the coelom, and metamerism, in which the body was built of repeated "modules" which could later specialize, such as the heads of most arthropods, which are composed of fused, specialized segments.
In comb jellies, there are species with one and sometimes two permanent anuses, species like the warty comb jelly grows an anus, which then disappear when it is no longer needed. [4]
In animals at least as complex as an earthworm, the embryo forms a dent on one side, the blastopore, which deepens to become the archenteron, the first phase in the growth of the gut. In deuterostomes, the original dent becomes the anus while the gut eventually tunnels through to make another opening, which forms the mouth. The protostomes were so named because it was thought that in their embryos the dent formed the mouth first ( proto– meaning "first") and the anus was formed later at the opening made by the other end of the gut. Research from 2001 shows the edges of the dent close up in the middles of protosomes, leaving openings at the ends which become the mouths and anuses. [5]
Bilateria is a large clade or infrakingdom of animals called bilaterians, characterized by bilateral symmetry during embryonic development. This means their body plans are laid around a longitudinal axis with a front and a rear end, as well as a left–right–symmetrical belly (ventral) and back (dorsal) surface. Nearly all bilaterians maintain a bilaterally symmetrical body as adults; the most notable exception is the echinoderms, which extend to pentaradial symmetry as adults, but are only bilaterally symmetrical as an embryo. Cephalization is also a characteristic feature among most bilaterians, where the special sense organs and central nerve ganglia become concentrated at the front/rostral end.
The gastrointestinal tract is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and other animals, including the esophagus, stomach, and intestines. Food taken in through the mouth is digested to extract nutrients and absorb energy, and the waste expelled at the anus as feces. Gastrointestinal is an adjective meaning of or pertaining to the stomach and intestines.
The coelom is the main body cavity in many animals and is positioned inside the body to surround and contain the digestive tract and other organs. In some animals, it is lined with mesothelium. In other animals, such as molluscs, it remains undifferentiated. In the past, and for practical purposes, coelom characteristics have been used to classify bilaterian animal phyla into informal groups.
Defecation follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus or cloaca. The act has a variety of names ranging from the common, like pooping or crapping, to the technical, e.g. bowel movement, to the obscene (shitting), to the euphemistic, to the juvenile. The topic, usually avoided in polite company, can become the basis for some potty humor.
A body orifice is any opening in the body of an animal.
A cloaca, pl.: cloacae, or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts of many vertebrate animals. All amphibians, reptiles, birds, and a few mammals have this orifice, from which they excrete both urine and feces; this is in contrast to most placental mammals, which have separate orifices for evacuation and reproduction. Excretory openings with analogous purpose in some invertebrates are also sometimes called cloacae. Mating through the cloaca is called cloacal copulation and cloacal kissing.
The archenteron, also called the gastrocoel, the primitive digestive tube or the primitive gut, is the internal cavity of the primitive gastrointestinal tract that forms during gastrulation in a developing animal embryo. It develops into the endoderm and mesoderm of the animal.
The cloaca is a structure in the development of the urinary and reproductive organs.
The mouth is the body orifice through which many animals ingest food and vocalize. The body cavity immediately behind the mouth opening, known as the oral cavity, is also the first part of the alimentary canal, which leads to the pharynx and the gullet. In tetrapod vertebrates, the mouth is bounded on the outside by the lips and cheeks — thus the oral cavity is also known as the buccal cavity — and contains the tongue on the inside. Except for some groups like birds and lissamphibians, vertebrates usually have teeth in their mouths, although some fish species have pharyngeal teeth instead of oral teeth.
The rectum is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. Before expulsion through the anus or cloaca, the rectum stores the feces temporarily. The adult human rectum is about 12 centimetres (4.7 in) long, and begins at the rectosigmoid junction at the level of the third sacral vertebra or the sacral promontory depending upon what definition is used. Its diameter is similar to that of the sigmoid colon at its commencement, but it is dilated near its termination, forming the rectal ampulla. It terminates at the level of the anorectal ring or the dentate line, again depending upon which definition is used. In humans, the rectum is followed by the anal canal, which is about 4 centimetres (1.6 in) long, before the gastrointestinal tract terminates at the anal verge. The word rectum comes from the Latin rēctumintestīnum, meaning straight intestine.
The urbilaterian is the hypothetical last common ancestor of the bilaterian clade, i.e., all animals having a bilateral symmetry.
In humans, the anus is the external opening of the rectum located inside the intergluteal cleft. Two sphincters control the exit of feces from the body during an act of defecation, which is the primary function of the anus. These are the internal anal sphincter and the external anal sphincter, which are circular muscles that normally maintain constriction of the orifice and which relax as required by normal physiological functioning. The inner sphincter is involuntary and the outer is voluntary. Above the anus is the perineum, which is also located beneath the vulva or scrotum.
Deuterostomes are bilaterian animals of the superphylum Deuterostomia, typically characterized by their anus forming before the mouth during embryonic development. Deuterostomia is further divided into four phyla: Chordata, Echinodermata, Hemichordata, and the extinct Vetulicolia known from Cambrian fossils. The extinct clade Cambroernida is thought to be a member of Deuterostomia.
Phoronids are a small phylum of marine animals that filter-feed with a lophophore, and build upright tubes of chitin to support and protect their soft bodies. They live in most of the oceans and seas, including the Arctic Ocean but excluding the Antarctic Ocean, and between the intertidal zone and about 400 meters down. Most adult phoronids are 2 cm long and about 1.5 mm wide, although the largest are 50 cm long.
Protostomia is the clade of animals once thought to be characterized by the formation of the organism's mouth before its anus during embryonic development. This nature has since been discovered to be extremely variable among Protostomia's members, although the reverse is typically true of its sister clade, Deuterostomia. Well-known examples of protostomes are arthropods, molluscs, annelids, flatworms and nematodes. They are also called schizocoelomates since schizocoely typically occurs in them.
The embryological origin of the mouth and anus is an important characteristic, and forms the morphological basis for separating bilaterian animals into two natural groupings: the protostomes and deuterostomes.
In evolutionary developmental biology, inversion refers to the hypothesis that during the course of animal evolution, the structures along the dorsoventral (DV) axis have taken on an orientation opposite that of the ancestral form.
The evolution of nervous systems dates back to the first development of nervous systems in animals. Neurons developed as specialized electrical signaling cells in multicellular animals, adapting the mechanism of action potentials present in motile single-celled and colonial eukaryotes. Primitive systems, like those found in protists, use chemical signalling for movement and sensitivity; data suggests these were precursors to modern neural cell types and their synapses. When some animals started living a mobile lifestyle and eating larger food particles externally, they developed ciliated epithelia, contractile muscles and coordinating & sensitive neurons for it in their outer layer.
Saccorhytus is an extinct genus of animal possibly belonging to the superphylum Ecdysozoa, and it is represented by a single species, Saccorhytus coronarius. The organism lived approximately 540 million years ago in the beginning of the Cambrian period. Initially proposed as a deuterostome, which would have made it the oldest known species of this superphylum, it has since been determined to belong to a protostome group called the ecdysozoans.
Xenoturbella bocki is a marine benthic worm-like species from the genus Xenoturbella. It is found in saltwater sea floor habitats off the coast of Europe, predominantly Sweden. It was the first species in the genus discovered. Initially it was collected by Swedish zoologist Sixten Bock in 1915, and described in 1949 by Swedish zoologist Einar Westblad. The unusual digestive structure of this species, in which a single opening is used to eat food and excrete waste, has led to considerable study and controversy as to its classification. It is a bottom-dwelling, burrowing carnivore that eats mollusks.