The Lacertidae are the family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at about 360 species in 39 genera. They represent the dominant group of reptiles found in Europe.
Anguidae refers to a large and diverse family of lizards native to the Northern Hemisphere. Common characteristics of this group include a reduced supratemporal arch, striations on the medial faces of tooth crowns, osteoderms, and a lateral fold in the skin of most taxa. The group is divided into two living subfamilies, the legless Anguinae, which contains slow worms and glass lizards, among others, found across the Northern Hemisphere, and Gerrhonotinae, which contains the alligator lizards, native to North and Central America. The family Diploglossidae was also formerly included. The family contains about 87 species in 8 genera.
Peltosaurus is an extinct genus of anguid lizard from North America that lived from the Eocene to the Oligocene. Peltosaurus belongs to the anguid subfamily Glyptosaurinae. The type species Peltosaurus granulosus was named in 1873 by American paleontologist Edward Drinker Cope. Many additional species have been named, but most have been reassigned to different genera. For example, Peltosaurus piger, named in 1928, was reclassified as Odaxosaurus piger, and P. jepseni, named in 1942 from the Paleocene of Wyoming, but was later reclassified as Proxestops jepseni. In 1955 a new species, Peltosaurus macrodon, was named from the Eocene of California. Lizard bones from the Late Miocene of Nebraska were attributed to a new species of Peltosaurus called P. minimus in 1976, extending the fossil range of Peltosaurus and Glyptosaurinae into the Neogene. However, these bones were later referred to a genus of skinks called Eumeces, meaning that the fossil range of Peltosaurus and Glyptosaurinae does not go beyond the Paleogene.
Blanus, also known as worm lizards, are a genus of amphisbaenians found in the Mediterranean region of Europe and North Africa. Like other amphisbaenians, Blanus species are specialized for a subterranean existence, with long, slender bodies, reduced limbs, and rudimentary eyes. Their skulls are powerfully constructed, allowing them to push through soil to create a burrow. Their jaws are well-developed, with large, recurved teeth and a pair of canine-like teeth in the upper jaw.
Headonhillia is an extinct genus of anguine lizard which existed in England during the Eocene epoch. It was first named by Jozef Klembara and Bryony Green in 2010, and represents one of the smallest anguines currently known. The type species is Headonhillia parva. Fossils have been found in the Hampshire Basin.
Saniwa is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species S. ensidens was described in 1870 as the first fossil lizard known from North America. A second species, S.orsmaelensis, is recognised from remains found in Europe. It is a close relative of Varanus, the genus that includes monitor lizards.
Bahndwivici is an extinct genus of lizard known from a nearly complete and articulated skeleton discovered in rocks of the Green River Formation of Wyoming, United States. The skeleton is very similar to that of the modern Chinese crocodile lizard, Shinisaurus.
Barbaturex is an extinct genus of giant herbivorous iguanian lizards from Pondaung Formation, the Eocene of Myanmar. It is represented by a single species, Barbaturex morrisoni, which is known from several partial dentaries and a fused pair of frontals, two bones that form part of the top of the skull. Based on the size of these bones, Barbaturex morrisoni is estimated to have been about 1 metre (3.3 ft) from snout to vent, and possibly up to 6 feet (1.8 m) including the tail. Barbaturex morrisoni was named after The Doors frontman Jim Morrison, a play on his epithet "The Lizard King". The genus's name is a portmanteau of the Latin words Barbatus and rex, meaning "bearded king", in reference to ridges along the mandible and the lizard's large size.
Shinisauria is a clade or evolutionary grouping of anguimorph lizards that includes the living Chinese crocodile lizard Shinisaurus and several of its closest extinct relatives. Shinisauria was named in 2008 by Jack Lee Conrad as a stem-based taxon to include all anguimorphs more closely related to Shinisaurus than to Anguis fragilis, Heloderma suspectum or Varanus varius. Several recent phylogenetic analyses of lizard evolutionary relationships place Shinisauria in a basal position within the clade Platynota, which also includes monitor lizards, helodermatids, and the extinct mosasaurs. Shinisaurians were once thought to be closely related to the genus Xenosaurus, but they are now considered distant relatives within Anguimorpha. The fossil record of shinisaurians extends back to the Early Cretaceous with Dalinghosaurus, which is from the Aptian aged Yixian Formation of China. Two other extinct shinisaurians are currently known: Bahndwivici from the Eocene of Wyoming and Merkurosaurus from the Late Oligocene of Germany and the Early Miocene of the Czech Republic. An indeterminate shinisaurian is known from an isolated tail found in the Eocene aged Messel pit in Germany.
Spathorhynchus is an extinct genus of amphisbaenians or worm lizards that existed from the Middle Eocene to the Early Oligocene in what is now Wyoming. It includes two species, the type species S. fossorium, named in 1973 from the Middle Eocene Bridger and Wind River Formations, and the species S. natronicus, named in 1977 from the Lower Oligocene White River Formation. Spathorhynchus belongs to the family Rhineuridae, which includes many other extinct species that ranged across North America at various times in the Cenozoic but only has one surviving member, Rhineura floridana, from Florida. Spathorhynchus differs from all other rhineurids except Dyticonastis from the Late Oligocene-Early Miocene of Oregon in having a slightly widened, spatula-shaped snout tip with a low angle of about 30 degrees. The two taxa may be closely related, having evolved in isolation in western North America after the formation of the Rocky Mountains separated them from rhineurids further east.
Ototriton is an extinct genus of rhineurid amphisbaenian or worm lizard from the Early Eocene of the western United States, including the type and only species Ototriton solidus. Paleontologist F. B. Loomis named Ototriton in 1919 on the basis of a single skull from the Wind River Formation in Wyoming, misinterpreting it as the skull of a salamander. Unlike salamanders and like other rhineurids, Ototriton has a shovel-shaped snout that it presumably used for burrowing underground. Ototriton is one of the earliest known rhineurids and also one of the largest.
Anniealexandria is an extinct genus of amphisbaenian lizard known by the type species Anniealexandria gansi from the earliest Eocene of Wyoming. Anniealexandria is the only known member of the family Bipedidae in the fossil record, which otherwise only includes the extant genus Bipes from Mexico. It was named in 2009 in honor of Annie Montague Alexander, founder of the University of California Museum of Paleontology. Remains of Anniealexandria are known only from a single fossil locality in the Bighorn Basin called Castle Gardens, but within the locality its fossils are common in the Willwood Formation, usually consisting of isolated jaw bones and vertebrae. Anniealexandria seems to have been a common component of a paleofauna that included fifteen other lizard species and existed in western North America during a period of global warming in the latest Paleocene and earliest Eocene.
Suzanniwana is an extinct genus of iguanian lizards that lived in western North America during the earliest Eocene, approximately 56 million years ago. Two species are known from the Bighorn Basin of Wyoming: the type species S. patriciana named in 2009, and the species S. revenanta named in 2013. Suzanniwana lived during the Paleocene–Eocene Thermal Maximum, a brief period of global warming that resulted in warmer and drier conditions in the Bighorn Basin. It likely stemmed from a lineage that had migrated into the basin from regions farther to the south, following a latitudinal band of constant climatic conditions that moved northward as the planet warmed. Suzanniwana shares many skeletal features with modern casquehead lizards of the family Corytophanidae and may be a stem-corytophanid. It also closely resembles Geiseltaliellus, an iguanian from the middle Eocene Messel pit in Germany.
Anolbanolis is an extinct genus of iguanian lizards that lived in the Bighorn Basin of what is now Wyoming during the earliest Eocene. The type species A. banalis was named by paleontologist Krister Smith in 2009 from a collection of isolated skull fragments found in a layer of the Willwood Formation that dates to a brief period of global warming called the Paleocene-Eocene thermal maximum (PETM) about 56 million years ago. The genus name Anolbanolis means "poor anole" from the Greek anolbos and the name Anolis, in reference to the scrappy nature of known fossil material and its close resemblance to lizards in the genus Anolis. Smith suggested that Anolbanolis may be a close relative of Anolis or Polychrus, which are common today in Central and South America but not found as far north as Wyoming, fitting with the idea that the Bighorn Basin was warmer and wetter in the Eocene than it is currently. According to Smith, the species name banalis is a reference to it being a "banal" lizard in the Willwood, being "quite abundant in the type locality," and not unusual because "the discovery of Eocene boreal fossil members of living subtropical and tropical clades is becoming commonplace." In 2011 Smith named a second species of Anolbanolis, A. geminus, which lived in the Bighorn Basin shortly after the PETM and was about 50 percent larger than A. banalis. Smith concluded on the basis of features in A. geminus that Anolbanolis is more closely related to Anolis than it is to Polychrus.
Babibasiliscus is an extinct genus of casquehead lizard that lived in what is now Wyoming during the early Eocene, approximately 48 million years ago. The genus is known from a single species, Babibasiliscus alxi, which was named by paleontologist Jack Conrad in 2015 on the basis of a fossilized skull from the Bridger Formation in the Green River Basin. The name Babibasiliscus comes from the Shoshoni word babi, meaning "older male cousin", and Basiliscus, a modern-day genus of casquehead lizards. The specimen is undeformed and nearly complete except for the tip of the snout and the top of the skull, making it unclear whether the distinctive bony crest of living corytophanids was present in prehistoric relatives like Babibasiliscus. The skull is about 42 millimetres (2 in) in length and the entire body is estimated to have been about 0.6 metres (2 ft) long. Bones on the right side of lower jaw of the specimen are thickened and fused together, suggesting that the jaw had broken and healed when the animal was alive.
Indiagama is an extinct genus of agamid lizard known from the type species Indiagama gujarata from the early Eocene of India. Indiagama was named in 2013 on the basis of a single lower jaw from the Cambay Shale in Gujarat. The rectangular shape of its teeth distinguish it from all other agamids, living and extinct.
Suratagama is an extinct genus of agamid lizard known from the type species Suratagama neeraae from the early Eocene of India. It was named in 2013 on the basis of three isolated jaw bones from the Cambay Shale in Gujarat.
Geiseltaliellus is an extinct genus of iguanian lizards that lived in what is now western Europe during the Eocene. It belongs to the family Corytophanidae, which includes modern casquehead lizards. Many fossils are known from Germany, France, and Belgium, with the most well preserved coming from the Messel pit lagerstätte in Messel, Germany. German paleontologist Oskar Kuhn named the genus in 1944 after the Geiseltal valley where the first specimens were found, designating the type species Geiseltaliellus longicaudus. Three new species — G. louisi, G. lamandini, and G. grisolli — were named in the 1990s and 2000s on the basis of more fragmentary remains from France and Belgium, although G. louisi has since been synonymized with G. longicaudus. In 2009 the Messel pit specimens were recognized as belonging to a species distinct from that of the G. longicaudus specimens in Geiseltalt and were collectively reclassified under a new name, G. maarius.
Paranecrosaurus is an extinct genus of lizard from the Eocene Messel Pits of Germany. It contains a single species, Paranecrosaurus feisti, originally described as a species of Saniwa. It was carnivorous, as indicated by the presence of the lizard Cryptolacerta as stomatch contents. It is placed in the family Palaeovaranidae.
Archaeovaranus is genus of varanid lizard from the early Eocene (Ypresian) Yuhuangding Formation of Hubei Province, China. The genus contains a single species, Archaeovaranus lii, known from a nearly complete skeleton. The holotype, which includes an intact skull, is associated but disarticulated. Archaeovaranus fills a gap in the varanid fossil record, as it represents a stem-varanid from the early Eocene of East Asia, and is the closest known relative of Varanus.
