Cambroproteus Temporal range: | |
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Juvenile and adult C. lemdadensis resting on Acadoparadoxides sp. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita |
Order: | † Ptychopariida |
Genus: | † Cambroproteus |
Species: | †C. lemdadensis |
Binomial name | |
†Cambroproteus lemdadensis | |
Cambroproteus is a genus of ptychopariid trilobite from the Late Cambrian of Morocco. [1] It has a single known species, Cambroproteus lemdadensis, whose various features are shared with disparate groups of other ptychopariids, which make it difficult to place within other established ptychopariid families. [1]
The generic name translates as "Cambrian Proteus." [1] This is in reference to the situation of how its varied features stymy understanding of its placement within Ptychopariida, effectively making it a marine shapeshifter. [1]
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Nektaspida is an extinct order of non-mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.
Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.
Agraulos is a genus of Solenopleuridae trilobites that lived during the Middle Cambrian in North America and Europe, particularly the Czech Republic. The genus was named by Hawle & Corda in 1847.
Palaeolenus is an extinct genus of ptychopariid trilobites of the family Palaeolenidae. Species lived during the later part of the Botomian stage of the Cambrian Period, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period
Lemdadella is an extinct genus of redlichiid trilobites that lived during the late Atdabanian stage, which lasted from 521 to 514 million years ago during the early part of the Cambrian Period.
Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.
The Wheeler Shale is a Cambrian (c. 507 Ma) fossil locality world-famous for prolific agnostid and Elrathia kingii trilobite remains and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna and preservation style normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten.
Trilobites are used as index fossils to subdivide the Cambrian period. Assemblages of trilobites define trilobite zones. The Olenellus-zone has traditionally marked the top of the Lower Cambrian, and is followed by the Eokochaspis zone.
Olenus is a genus of Upper Cambrian ptychopariid trilobite.
The Olenelloidea are a superfamily of trilobites, a group of extinct marine arthropods. They lived during the late Lower Cambrian and species occurred on all paleocontinents.
Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
The Terreneuvian is the lowermost and oldest series of the Cambrian geological system. Its base is defined by the first appearance datum of the trace fossil Treptichnus pedum around 538.8 million years ago. Its top is defined as the first appearance of trilobites in the stratigraphic record around 521 million years ago. This series' name was formally accepted by the International Commission on Stratigraphy in 2007.
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about 521 million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera Olenellus or Redlichia around 514 million years ago.
Agnostotes orientalis is a species of agnostid trilobite belonging to the genus Agnostotes. It existed during the Jiangshanian Age of the Cambrian. It is an important index fossil in biostratigraphy.
Cambrian Series 2 is the unnamed 2nd series of the Cambrian. It lies above the Terreneuvian series and below the Miaolingian. Series 2 has not been formally defined by the International Commission on Stratigraphy, lacking a precise lower boundary and subdivision into stages. The proposed lower boundary is the first appearance of trilobites which is estimated to be around 521 million years ago.
Condylopyge Hawle and Corda (1847) is a genus of agnostid trilobite that lived during the late Lower and early Middle Cambrian, in what are today Canada, the Czech Republic, England and Wales, France, Germany, Italy, Morocco, the Russian Federation, Spain, Turkey and Sweden. It can easily be distinguished from all other Agnostida because the frontal glabellar lobe is notably wider than the rear lobe. It belongs to the same family as Pleuroctenium but the frontal glabellar lobe does not fold around the rear lobe, as it does in that genus. Condylopyge is long ranging, possibly spanning the early Cambrian Terreneuvian Series in Nuneaton, central England into at least Drumian strata at various locations elsewhere.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.
Galeaspis is an extinct genus of ptychopariid trilobite in the family Catillicephalidae. Species are found in the Seletinian Horizon of the Kuyandinian faunal stage of Late Cambrian marine strata in Central Kazakhstan. Fossil specimens are usually semicircular to bean-shaped, incomplete cranidia, with globular glabella.