Cestites

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Cestites
Cestites mirabilis holotype.jpg
Holotype of Cestites mirabilis, from Douglas Lake Member of Lenoir Limestone, at Douglas Dam, Tennessee [1]
Scientific classification
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Cestites

Caster and Brooks (1956) [2]
Type species
Cestites mirabilis
Caster & Brooks (1956)

Cestites ("girdle") is a controversial fossil from the Middle Ordovician (Darriwilian, 460 million years old) Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee. [1] At first considered a ctenophore by Caster and Brooks, [2] it was later interpreted as a genus of liverwort by Gregory Retallack.

Contents

Thallus of Cestites mirabilis with archegoniophores Cestites mirabilis fertile.jpg
Thallus of Cestites mirabilis with archegoniophores
Interpretaive sketch of Cestites mirabilis Cestites mirabilis sketch.tif
Interpretaive sketch of Cestites mirabilis
Spore of Cestites mirabilis Cestites mirabilis spore.tif
Spore of Cestites mirabilis

Description

Cestites has a narrow gametophyte thallus, with a wide midrib and dichotomizing at long intervals. The archegoniophores are parasol shaped and clustered.

Interpretation

The liverwort interpretation of this fossil has been considered controversial in some quarters [3] but accepted elsewhere. [4]

Related Research Articles

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The Cambrian Period is the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period 485.4 mya. Its subdivisions, and its base, are somewhat in flux.

<span class="mw-page-title-main">Silurian</span> Third period of the Paleozoic Era, 443–419 million years ago

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<span class="mw-page-title-main">Marchantiophyta</span> Botanical division of non-vascular land plants

The Marchantiophyta are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information.

<span class="mw-page-title-main">Hornwort</span> Division of non-vascular land plants with horn-shaped sporophytes

Hornworts are a group of non-vascular Embryophytes constituting the division Anthocerotophyta. The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information; the flattened, green plant body of a hornwort is the gametophyte stage of the plant.

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<span class="mw-page-title-main">Chasmataspidida</span> Order of arthropods

Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hind part (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural group.

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Polysporangiophytes, also called polysporangiates or formally Polysporangiophyta, are plants in which the spore-bearing generation (sporophyte) has branching stems (axes) that bear sporangia. The name literally means 'many sporangia plant'. The clade includes all land plants (embryophytes) except for the bryophytes whose sporophytes are normally unbranched, even if a few exceptional cases occur. While the definition is independent of the presence of vascular tissue, all living polysporangiophytes also have vascular tissue, i.e., are vascular plants or tracheophytes. Extinct polysporangiophytes are known that have no vascular tissue and so are not tracheophytes.

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<span class="mw-page-title-main">Gregory Retallack</span> American paleontologist

Gregory John Retallack is an Australian paleontologist, geologist, and author who specializes in the study of fossil soils (paleopedology). His research has examined the fossil record of soils though major events in Earth history, extending back some 4.6 billion years. Among his publications he has written two standard paleopedology textbooks, said N. Jones in Nature Geoscience "Retallack has literally written the book on ancient soils."

<span class="mw-page-title-main">Douglas Lake Member</span> Geologic formation in [[Tennessee]], United States

The Douglas Lake Member is a geologic unit of member rank of the Lenoir Limestone that overlies the Mascot Dolomite and underlies typical nodular member of the Lenoir Limestone in Douglas Lake, Tennessee, region. It fills depressions that are part of a regional unconformity at the base of Middle Ordovician strata, locally the Lenoir Limestone, that separates them from the underlying Lower Ordovician strata, locally the Knox Group.

<i>Casterlorum</i> Genus of hornwort fossil

Casterlorum is a genus of fossil with controversial interpretation from the Middle Ordovician Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee The genus was named in honor of Ken Caster.

<i>Janegraya</i> Extinct genus of liverworts

Janegraya is a genus of fossil with controversial interpretation from the Middle Ordovician Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee. The generic name honors Jane Gray, and the epithet means "prophetess".

<i>Dollyphyton</i> Extinct genus of mosses

Dollyphyton is a genus of fossil with controversial interpretation from the Middle Ordovician Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee. The generic name honors Dolly Parton whose Dollywood resort is nearby. The epithet honors Art Boucot.

<i>Edwardsiphyton</i> Extinct genus of mosses

Edwardsiphyton is a genus of fossil from the Middle Ordovician Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee The genus was named in honor of Dianne Edwards, and the epithet refers to the shape of the capsules.

<i>Chasmataspis</i> Extinct genus of arthropods

Chasmataspis is a genus of chasmataspidid, a group of extinct aquatic chelicerate arthropods. It was found in the Early Ordovician deposits of Tennessee, United States.

<i>Palaeoglomus</i> Genus of fungi

Palaeoglomus is a genus of microscopic mycorrhizal fossil, found in palynological preparations of rocks which separate out organic remains by acid dissolution.

<i>Douglasocaris</i> Genus of small freshwater animals

Douglasocaris is a genus of bivalved arthropod from the Middle Ordovician Douglas Lake Member of the Lenoir Limestone from Douglas Dam Tennessee.

References

  1. 1 2 Retallack, G.J. (2019). "Ordovician land plants and fungi from Douglas Dam, Tennessee". The Palaeobotanist. 68: 1–33.
  2. 1 2 Caster, K.E.; Brooks, H.K. (1956). "New fossils from the Canadian–Chazyan (Ordovician) hiatus in Tennessee". Bulletins of American Paleontology. 36: 157–199.
  3. Edwards, Dianne; Morris, Jennifer L.; Axe, Lindsey; Duckett, Jeffrey G.; Pressel, Silvia; Kenrick, Paul (2022). "Piecing together the eophytes – a new group of ancient plants containing cryptospores". New Phytologist. 233 (3): 1440–1455. doi: 10.1111/nph.17703 . ISSN   0028-646X. PMID   34806774. S2CID   244495761.
  4. Leigh, Egbert (2022). "Fossil soils: trace fossils of ecosystems on land and windows on the context of evolution". Evolution Education and Outreach. 15:14: 1–5. doi: 10.1186/s12052-022-00173-3 . S2CID   252203727.