Chalicotherioidea Temporal range: Eocene to Middle Pleistocene Early | |
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Moropus elatus at the National Museum of Natural History, Washington, DC | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Infraorder: | † Ancylopoda |
Superfamily: | † Chalicotherioidea Gill, 1872 |
Families | |
Chalicotherioidea is an extinct superfamily of clawed perissodactyls (odd-toed ungulates) that lived from the early Eocene to the early Pleistocene subepochs. [1] Based on the fossil record they emerged and thrived largely in Eurasia, although specimens have been found in both Africa and North America. They were likely browsers that fed mainly on leaves, twigs, and other nonresistant vegetation. Many of the contained genera had derived specializations of the forelimb and manus that allowed the claws to be used as hooks for browsing and to be kept off of the ground while walking. [2] Chalicotheres lived primarily in forested areas. Size sexual dimorphism and morphological structures such as the domed skulls of Tylocephalonyx suggest agonistic behaviour in some sort of social setting. [2] They are related to modern day horses, rhinoceroses, and tapirs, as well as the extinct brontotheres. [3]
Ancylopoda is a group of browsing, herbivorous, mammals in the Perissodactyla that show long, curved and cleft claws. Morphological evidence indicates the Ancylopoda diverged from the tapirs, rhinoceroses and horses (Euperissodactyla) after the Brontotheria; however, earlier authorities such as Osborn sometimes considered the Ancylopoda to be outside Perissodactyla or, as was popular more recently, to be related to Brontotheriidae.
Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla, are animals—ungulates—who have reduced the weight-bearing toes to three or one of the five original toes. The non-weight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, the even-toed ungulates bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines, rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do.
Ungulates are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. Living ungulates are divided into two orders: the odd-toed ungulates (Perissodactyla) including horses, rhinoceroses, and tapirs; and even-toed ungulates (Artiodactyla) such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
The even-toed ungulates are ungulates—hoofed animals—which bear weight equally on two of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many other even-toed ungulates digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do.
Creodonta is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the Oxyaenids and the Hyenodonts, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa.
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates. It is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Artiodactyla, being composed thus of several unrelated lineages.
Chalicotheres are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 mya. They are one of the five major radiations of perissodactyls, with three groups living, and two extinct.
Phenacodus is an extinct genus of mammals from the late Paleocene through middle Eocene, about 55 million years ago. It is one of the earliest and most primitive of the ungulates, typifying the family Phenacodontidae and the order Perissodactyla.
Chalicotherium is a genus of extinct odd-toed ungulates of the order Perissodactyla and family Chalicotheriidae. The genus is known from Europe and Asia, from the Middle Miocene to Late Miocene.
Eomoropidae is a family of odd-toed ungulates, a group which also includes horses, rhinoceroses, and tapirs. They were most closely related to the extinct chalicotheres, which they greatly resemble, and may have been their immediate ancestors. They were, however, much smaller than the later forms, being around the size of a sheep. Like their later relatives, they were probably browsers on leaves and other soft vegetation, and, unlike most other ungulates, had claws on their feet. The Eomoropidae is most likely a paraphyletic group within Chalicotherioidea.
Palaearctonyx is an extinct genus of omnivorous placental mammals from clade Carnivoraformes, that lived in North America from early to middle Eocene.
Bryanictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America, from the early to late Paleocene.
Intyrictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America during early Paleocene.
Pristinictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America during middle Paleocene.
Cormocyon is an extinct genus of borophagine canid native to North America. It lived from the Oligocene to the Early Miocene, 30.8—20.6 Mya, existing for about 10.2 million years. It is regarded as a primitive, transitional member of the Borophagini tribe.
Protepicyon is an extinct monospecific genus of the Borophaginae subfamily of canids native to North America. It lived during the Barstovian stage of the Middle Miocene 16.0—13.6 mya. One of the top predators of its time, it was the probable ancestor of the better known Epicyon, and is known from remains in California and New Mexico.
Raphictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America during late Paleocene.
Hypohippus is an extinct genus of three-toed horse, which lived 17–11 million years ago. It was the largest anchitherine equid about the size of a modern domestic horse, at 403–600 kg (888–1,323 lb) and 1.8 m (5.9 ft) long. It was a long-necked, high-shouldered browser with sub-hypsodont, lophodont (rhino-like) dentition, that fed on the tough vegetation of forest understory and shrubs. Its deep preorbital fossae and retraction of the nasal notch hint at the presence of a long, muscular and prehensile upper lip that would aid during selective browsing. Overall its ecology would have been more comparable to modern okapi than to grazing horses. Fossils of it have been found in Nebraska, Colorado, and Montana.
Hyraxes, also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between 30 and 70 cm long and weigh between 2 and 5 kg. They are superficially similar to pikas and marmots, but are more closely related to elephants and sea cows.
Barylambdidae is an extinct family of pantodont mammals from North America.