Dicharax

Dicharax
Shell of Dicharax abdoui (holotype at MNHN, Paris)
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Architaenioglossa
Superfamily:	Cyclophoroidea
Family:	Alycaeidae
Genus:	Dicharax Kobelt & Möllendorff, 1900
Type species
Alycaeus hebesBenson, 1857
Synonyms
List Alycaeus (Charax) Benson, 1859 Alycaeus (Dicharax) Kobelt & Möllendorff, 1900 AwalycaeusKuroda, 1951 Chamalycaeus (Cipangocharax) Kuroda, 1943 Chamalycaeus (Dicharax) Kobelt & Möllendorff, 1900 Chamalycaeus (Sigmacharax) Kuroda, 1943 CipangocharaxKuroda, 1943

Dicharax is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Alycaeidae. ^[1]

Species

Species within the genus Dicharax include:

• Dicharax abdoui Páll-Gergely, 2017
• Dicharax abei (Kuroda, 1951)
• Dicharax admirandus Páll-Gergely & Auffenberg, 2021
• Dicharax akhaensis (Godwin-Austen, 1914)
• Dicharax akioi (Kuroda & Abe, 1980)
• Dicharax akiratadai (Minato, 1982)
• Dicharax alticola Páll-Gergely & Hunyadi, 2017
• Dicharax ananensis (Yano, Akira & H. Matsuda, 2013)
• Dicharax anonymus (Godwin-Austen, 1914)
• Dicharax anthostoma (Möllendorff, 1885)
• Dicharax asaluensis (Godwin-Austen, 1914)
• Dicharax ataranensis (Godwin-Austen, 1914)
• Dicharax avae (W. T. Blanford, 1863)
• Dicharax barowliensis (Godwin-Austen, 1914)
• Dicharax bawai Aravind & Páll-Gergely, 2018
• Dicharax bicrenatus (Godwin-Austen, 1874)
• Dicharax biexcisus (Pilsbry, 1902)
• Dicharax bifrons (Theobald, 1870)
• Dicharax birugosus (Godwin-Austen, 1893)
• Dicharax bison Páll-Gergely & Hunyadi, 2017
• Dicharax blanfordi (Godwin-Austen, 1914)
• Dicharax borealis Jirapatrasilp & Páll-Gergely, 2021
• Dicharax burchi Jirapatrasilp & Páll-Gergely, 2021
• Dicharax burroiensis (Godwin-Austen, 1914)
• Dicharax candrakirana Nurinsiyah & Hausdorf, 2017
• Dicharax caudapiscis Páll-Gergely & Hunyadi, 2018
• Dicharax chennelli (Godwin-Austen, 1886)
• Dicharax conicus (Godwin-Austen, 1871)
• Dicharax crassicollis (van Benthem Jutting, 1959)
• Dicharax crenatus (Godwin-Austen, 1871)
• Dicharax crispatus (Godwin-Austen, 1871)
• Dicharax cristatus (Möllendorff, 1886)
• Dicharax cucullatus (Theobald, 1870)
• Dicharax cyclophoroides (Pilsbry & Y. Hirase, 1909)
• Dicharax cyclostoma Páll-Gergely, 2021
• Dicharax daflaensis (Godwin-Austen, 1876)
• Dicharax dalingensis (Godwin-Austen, 1914)
• Dicharax damsangensis (Godwin-Austen, 1886)
• Dicharax depressus (Bavay & Dautzenberg, 1912)
• Dicharax diagonius (Godwin-Austen, 1871)
• Dicharax digitatus (H. F. Blanford, 1871)
• Dicharax diminutus (Heude, 1885)
• Dicharax diplochilus (Möllendorff, 1887)
• Dicharax dohertyi (Godwin-Austen, 1893)
• Dicharax dolichodeiros (Heude, 1890)
• Dicharax draco Páll-Gergely & Hunyadi, 2017
• Dicharax duoculmen (Godwin-Austen, 1914)
• Dicharax duorugosus (Godwin-Austen, 1914)
• Dicharax ebrius Páll-Gergely & Auffenberg, 2021
• Dicharax edei (Godwin-Austen, 1914)
• Dicharax elevatus (Heude, 1886)
• Dicharax ellipticus Páll-Gergely, 2017
• Dicharax expanstoma (Minato, 1982)
• Dicharax expatriatus (W. T. Blanford & H. F. Blanford, 1860)
• Dicharax fargesianus (Heude, 1885)
• Dicharax fimbriatus (Bavay & Dautzenberg, 1912)
• Dicharax floridus Páll-Gergely & Hunyadi, 2022
• Dicharax footei (W. T. Blanford & H. F. Blanford, 1861)
• Dicharax fraterculus (Bavay & Dautzenberg, 1900)
• Dicharax gemma (Godwin-Austen, 1914)
• Dicharax gemmula (Benson, 1859)
• Dicharax generosus (Godwin-Austen, 1914)
• Dicharax glaber (W. T. Blanford, 1865)
• Dicharax globulus (Godwin-Austen, 1874)
• Dicharax gnomus Páll-Gergely, 2021
• Dicharax habiangensis (Godwin-Austen, 1914)
• Dicharax hebes (Benson, 1857)
• Dicharax humilis (W. T. Blanford, 1862)
• Dicharax imitator Páll-Gergely & Hunyadi, 2017
• Dicharax immaculatus Páll-Gergely, 2017
• Dicharax ingrami (W. T. Blanford, 1862)
• Dicharax itonis (Kuroda, 1943)
• Dicharax jaintiacus (Godwin-Austen, 1871)
• Dicharax japonicus (Martens, 1865)
• Dicharax kezamaensis (Godwin-Austen, 1914)
• Dicharax khasiacus (Godwin-Austen, 1871)
• Dicharax kiuchii (Minato & Abe, 1982)
• Dicharax floridus Páll-Gergely & Hunyadi, 2022
• Dicharax kurodatokubeii (Minato, 1987)
• Dicharax kurzianus (Theobald & Stoliczka, 1872)
• Dicharax lahupaensis (Godwin-Austen, 1914)
• Dicharax lectus (Godwin-Austen, 1914)
• Dicharax lenticulus (Godwin-Austen, 1874)
• Dicharax levis (Godwin-Austen, 1914)
• Dicharax logtakensis (Godwin-Austen, 1914)
• Dicharax longituba (Martens, 1864)
• Dicharax magnus (Godwin-Austen, 1893)
• Dicharax maosmaiensis (Godwin-Austen, 1922)
• Dicharax microcostatus Páll-Gergely, 2017
• Dicharax microdiscus (Möllendorff, 1887)
• Dicharax micropolitus Páll-Gergely & Hunyadi, 2017
• Dicharax mirounga Páll-Gergely, 2021
• Dicharax miyazakii (Takahashi & Habe, 1973)
• Dicharax moellendorffi (Kobelt & Möllendorff, 1897)
• Dicharax montanus G. Nevill, 1881
• Dicharax multirugosus (Godwin-Austen, 1874)
• Dicharax muspratti (Godwin-Austen, 1914)
• Dicharax mutatus (Godwin-Austen, 1876)
• Dicharax nagaensis (Godwin-Austen, 1871)
• Dicharax nakashimai (Minato, 1987)
• Dicharax nattoungensis (Godwin-Austen, 1914)
• Dicharax nishii (Minato, 2005)
• Dicharax nitidus (W. T. Blanford, 1862)
• Dicharax nongtungensis (Godwin-Austen, 1914)
• Dicharax notatus (Godwin-Austen, 1876)
• Dicharax notus (Godwin-Austen, 1914)
• Dicharax nowgongensis (Godwin-Austen, 1914)
• Dicharax obscurus (Godwin-Austen, 1914)
• Dicharax ochraceus (Godwin-Austen, 1893)
• Dicharax okamurai (Azuma, 1980)
• Dicharax okinawaensis (K. Uozumi, Yamamoto & Habe, 1979)
• Dicharax oligopleuris (Möllendorff, 1887)
• Dicharax omissus (Godwin-Austen, 1914)
• Dicharax oshimanus (Pilsbry & Y. Hirase, 1904)
• Dicharax ovatus Páll-Gergely & Auffenberg, 2021
• Dicharax ovatus Páll-Gergely & Auffenberg, 2021
• Dicharax pachitaensis (Godwin-Austen, 1886)
• Dicharax panhai Jirapatrasilp & Páll-Gergely, 2021
• Dicharax panshiensis (Chen, 1989)
• Dicharax parvulus (Möllendorff, 1887)
• Dicharax peilei (Preston, 1914)
• Dicharax placenovitas (Minato, 1981)
• Dicharax planorbulus (Heude, 1885)
• Dicharax plectocheilus (Benson, 1859)
• Dicharax plicilabris (Möllendorff, 1886)
• Dicharax politus (W. T. Blanford, 1865)
• Dicharax pongrati Jirapatrasilp & Tongkerd, 2021
• Dicharax pratatensis (Panha & Burch, 1997)
• Dicharax purus (Pilsbry & Y. Hirase, 1904)
• Dicharax pusillus (Godwin-Austen, 1871)
• Dicharax rechilaensis (Godwin-Austen, 1914)
• Dicharax robustus Páll-Gergely & Hunyadi, 2017
• Dicharax sandowayensis (Godwin-Austen, 1914)
• Dicharax sculpturus (Godwin-Austen, 1875)
• Dicharax serratus (Godwin-Austen, 1874)
• Dicharax shiibaensis (Minato, 2005)
• Dicharax shiosakimasahiroi (Yano, H. Matsuda & Nishi, 2016)
• Dicharax sonlaensis (Raheem & S. Schneider, 2017)
• Dicharax spatiosus Páll-Gergely & Hunyadi, 2022
• Dicharax spiracellum (Adams & Reeve, 1850)
• Dicharax stoliczkii (Godwin-Austen, 1874)
• Dicharax strangulatus (L. Pfeiffer, 1846)
• Dicharax strigatus (Godwin-Austen, 1874)
• Dicharax stuparum Páll-Gergely & Hunyadi, 2018
• Dicharax subculmen (Godwin-Austen, 1893)
• Dicharax subhumilis (Möllendorff, 1897)
• Dicharax subroseus Páll-Gergely, 2021
• Dicharax succineus (W. T. Blanford, 1862)
• Dicharax sylheticus (Godwin-Austen, 1914)
• Dicharax tadai (Kuroda & Kawamoto, 1956)
• Dicharax takahashii (Habe, 1976)
• Dicharax tanegashimae (Pilsbry, 1902)
• Dicharax tangmaiensis (D.-N. Chen & G.-Q. Zhang, 2001)
• Dicharax theobaldi (W. T. Blanford, 1862)
• Dicharax tokunoshimanus (Pilsbry & Y. Hirase, 1904)
• Dicharax tsushimanus (Pilsbry & Y. Hirase, 1909)
• Dicharax umbonalis (Benson, 1856)
• Dicharax verrucosus Páll-Gergely & Hunyadi, 2021
• Dicharax vestitus (W. T. Blanford, 1862)
• Dicharax woodthorpi (Godwin-Austen, 1914)
• Dicharax yanoshigehumii (Minato, 1987)
• Dicharax yanoshokoae (Yano & H. Matsuda, 2016)

Species brought into synonymy

• Dicharax canaliculatus (Möllendorff, 1894): synonym of Chamalycaeus canaliculatus (Möllendorff, 1894) (unaccepted combination)
• Dicharax crenulatus (Benson, 1859): synonym of Metalycaeus crenulatus (Benson, 1859) (unaccepted combination)
• Dicharax davisi (Godwin-Austen, 1914): synonym of Dicharax cucullatus (Theobald, 1870) (junior synonym)
• Dicharax neglectus (Heude, 1885): synonym of Metalycaeus rathouisianus (Heude, 1882) (unaccepted combination)
• Dicharax neglectus (Godwin-Austen, 1914): synonym of Metalycaeus godwinausteni Páll-Gergely, 2020 (invalid: senior homonym of Alycaeus neglectus Heude, 1885)
• Dicharax prosectus (Benson, 1857): synonym of Metalycaeus prosectus (Benson, 1857) (unaccepted combination)
• Dicharax rugosus (Godwin-Austen, 1914): synonym of Metalycaeus rugosus (Godwin-Austen, 1914) (unaccepted combination)
• Dicharax stylifer (Benson, 1857): synonym of Metalycaeus stylifer (Benson, 1857) (unaccepted combination)
• Dicharax teriaensis (Godwin-Austen, 1914): synonym of Metalycaeus teriaensis (Godwin-Austen, 1914) (unaccepted combination)
• Dicharax toruputuensis (Godwin-Austen, 1914): synonym of Metalycaeus toruputuensis (Godwin-Austen, 1914) (unaccepted combination)

References

1. MolluscaBase eds. (2021). MolluscaBase. Dicharax Kobelt & Möllendorff, 1900. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=1039734 on 2021-04-24

• Kuroda, T. (1943). New Land Shells from Tyugoku District (1) : A New Species belonging to a New Subgenus of the Genus Chamalycaeus. Venus (Japanese Journal of Malacology). 13(1-4): 7-11.
• Kuroda, T. (1951). On Awalycaeus abei, gen. et sp. nov. (Cyclophoridae)(The Celebration Number of Mr. T. Kanamaru's 60th Birthday). Venus (Japanese Journal of Malacology). 17(3): 73-74.

External links

• Benson, W. H. (1859). A sectional distribution of the genus Alycæus, Gray, with characters of six new species and of other Cyclostomidæ collected at Darjiling by W. T. Blanford, Esq., Geol. Survey. The Annals and Magazine of Natural History, Ser. 3. 3: 176–184
• Kobelt, W. & Möllendorff, O. von. (1900). Zur Systematik der Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft. 32(11-12): 186
• Páll-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. & Ablett, J.D. (2020). Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalog. ZooKeys. 981: 1–220
• Gude, G. K. (1921). Mollusca. III. Land Operculates. Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae.). In: Shipley, A. E. & Marshall, G. A. K. (eds.):. The Fauna of British India, including Ceylon and Burma. I–XIV, 386 pp. London (Taylor & Francis).