A squid is a mollusc with an elongated soft body, large eyes, eight arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida. Like all other cephalopods, squid have a distinct head, bilateral symmetry, and a mantle. They are mainly soft-bodied, like octopuses, but have a small internal skeleton in the form of a rod-like gladius or pen, made of chitin.
Myopsida is one of the four orders of squid. It consists of two families: the monotypic Australiteuthis and the diverse and commercially important Loliginidae. Some taxonomists classify this taxon as a suborder of the order Teuthida, in which case it is known as Myopsina. This reclassification is due to Myopsina and Oegopsina not being demonstrated to form a clade.
Belemnoids are an extinct group of marine cephalopod, very similar in many ways to the modern squid. Like them, the belemnoids possessed an ink sac, but, unlike the squid, they possessed ten arms of roughly equal length, and no tentacles. The name "belemnoid" comes from the Greek word βέλεμνον, belemnon meaning "a dart or arrow" and the Greek word είδος, eidos meaning "form".
Helicoprion is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of Helicoprion are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian to the Roadian stage of the Guadalupian. The closest living relatives of Helicoprion are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of Helicoprion via morphometric analysis of the tooth whorls found only H. davisii, H. bessonowi and H. ergassaminon to be valid, with some of the larger tooth whorls being outliers.
Platecarpus is an extinct genus of aquatic lizards belonging to the mosasaur family, living around 84–81 million years ago during the middle Santonian to early Campanian, of the Late Cretaceous period. Fossils have been found in the United States and possible specimens in Belgium and Africa. A well-preserved specimen of Platecarpus shows that it fed on moderate-sized fish, and it has been hypothesized to have fed on squid, and ammonites as well. Like other mosasaurs, it was initially thought to have swum in an eel-like fashion, although another study suggests that it swam more like modern sharks. An exceptionally well-preserved specimen of P. tympaniticus known as LACM 128319 shows skin impressions, pigments around the nostrils, bronchial tubes, and the presence of a high-profile tail fluke, showing that it and other mosasaurs did not necessarily have an eel-like swimming method, but were more powerful, fast swimmers. It is held in the Natural History Museum of Los Angeles County. Isotopic analysis on teeth specimens has suggested that this genus and Clidastes may have entered freshwater occasionally, just like modern sea snakes.
Palaeoctopus is an extinct genus of octopuses that lived during the Late Cretaceous. It contains one valid species, P. newboldi, which has been found in Lebanon.
Tusoteuthis is an extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although often called a squid, it is now thought to be more closely related to modern octopuses. Although many specimens from Western Interior Seaway of North America were described, at 2019, only one, poorly preserved specimen from Kansas is recognized as Tusoteuthis. One species, T. longa, is traditionally recognized. In 2019, due to poor preservation of holotype specimen, Tusoteuthis was considered likely to be a nomen dubium, and it was proposed that later described species are better to included in genus Enchoteuthis instead.
Cimolichthys is an extinct genus of large predatory marine aulopiform ray-finned fish known worldwide from the Late Cretaceous. It is the only member of the family Cimolichthyidae.
Belemnotheutis is an extinct coleoid cephalopod genus from the middle and upper Jurassic, related to but morphologically distinct from belemnites. Belemnotheutis fossils are some of the best preserved among coleoids. Remains of soft tissue are well-documented in some specimens, even down to microscopic muscle tissue. In 2008, a group of paleontologists even recovered viable ink from ink sacs found in several specimens.
Cheirothrix is an extinct genus of marine ray-finned fish from the Late Cretaceous. It is named for its enlarged pectoral fins, which show close convergence with modern flying fish.
Charitosomus is an extinct genus of prehistoric marine ray-finned fish from the Late Cretaceous, related to modern beaked salmons. They were nektonic carnivores in life.
The Platyrhinidae are a family of rays, commonly known as thornbacks due to their dorsal rows of large thorns. They resemble guitarfishes in shape. Though traditionally classified with stingrays, molecular evidence suggests they are more closely related to electric rays in the order Torpediniformes.
The paleontological sites of Lebanon contain deposits of well preserved fossils and include some species found nowhere else. Notable among these is the Lebanese lagerstätten of the Late Cretaceous age, which contain a well-preserved variety of different fossils. Some fossils date back to the Jurassic period, and younger fossils of mammals from a different site belong to the Miocene through the Pleistocene.
Belemnitida is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip: the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than 5 mm (0.20 in), though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey.
Cephalopod fins, sometimes known as wings, are paired flap-like locomotory appendages. They are found in ten-limbed cephalopods as well as in the eight-limbed cirrate octopuses and vampire squid. Many extinct cephalopod groups also possessed fins. Nautiluses and the more familiar incirrate octopuses lack swimming fins. An extreme development of the cephalopod fin is seen in the bigfin squid of the family Magnapinnidae.
The Sannine Formation, also called the Sannine Limestone, is a Cretaceous geologic formation in Lebanon. It is a Konservat-Lagerstätte that contains a high diversity of well-preserved fish, reptiles, and invertebrates from the Tethys Ocean within its three main localities: Haqel, Hjoula, and Nammoura.
Mimodactylus is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to the Sannine Formation. The owner of the quarry allowed the specimen to be prepared and scientifically described by an international team of researchers. When it was eventually sold, the buyer donated it to the MIM Museum in Beirut. In 2019, the researchers named the new genus and species Mimodactylus libanensis; the generic name refers to the MIM Museum, combined with the Greek word daktylos for "digit", and the specific name refers to Lebanon. The well-preserved holotype specimen is the first complete pterosaur from the Afro-Arabian continent, and the third pterosaur fossil known from Lebanon.
Muensterelloidea is a superfamily of stem-octopod cephalopods from the Early Jurassic to Late Cretaceous. Two families are currently identified, Muensterellidae, and Patelloctopodidae. The clade is the ancestral group from which modern octopus arose.
Enchoteuthis is an extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although it and its relative Tusoteuthis are often compared to squid, both are now thought to be more closely related to modern octopuses. Examination of gladius remains initially yielded an estimated mantle length about 2 m based on specimen once described as Tusoteuthis longa, close to or equal to that of the modern giant squid, although reclassification of this genus as a muensterelloid results in a much shorter total length, about 3 m (10 ft). Three species are currently recognized as valid: E. melanae, E. tonii, and E. cobbani.
Sahel Alma or Sahel Aalma is a Late Cretaceous paleontological site and Konservat-Lagerstätte in Keserwan-Jbeil, Lebanon. Located near the town of the same name, it documents well-preserved deepwater marine fossils dating to the late Santonian stage of the Cretaceous. It is often associated with the slightly older, similarly famous Sannine Formation sites, with these four sites being together referred to as the "Fish Beds" of Lebanon.
