Hay-scented buckler-fern | |
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Dryopteris aemula in Galloway, Southern Scotland | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Division: | Polypodiophyta |
Class: | Polypodiopsida |
Order: | Polypodiales |
Suborder: | Polypodiineae |
Family: | Dryopteridaceae |
Genus: | Dryopteris |
Species: | D. aemula |
Binomial name | |
Dryopteris aemula | |
Dryopteris aemula, the hay-scented buckler-fern [1] or hay-scented fern, is a species of perennial leptosporangiate fern.
Dryopteris aemula grows as a crown of fronds arising from a short ascending rhizome.
The rachis is dark purple-brown with red-brown lanceolate scales. Leaves are tri-pinnate, triangular-ovate or triangular-lanceolate, 15–60 cm (5.9–23.6 in) long, often arching, semi-evergreen and pale yellow-green. Scattered small sessile glands grow on the underside or both surfaces of the fronds. Pinnae are slightly concave giving the frond a characteristic crisped appearance. The plant is hay-scented. [2]
Sori or spore-producing organs occur in a row down each side of the midrib. The indusium of the sorus is irregularly toothed and edged with sessile glands.
The plant does not reproduce vegetatively and relies upon spores to generate new individuals. [3]
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Dryopteris aemula is confined to the atlantic coastal areas of western Europe and Macaronesia. D. aemula is highly oceanic in its distribution occurring mostly in the Köppen climate type Cfb (oceanic climates). Even within Great Britain, it is restricted to western districts and has just a few outlying eastern localities such as The Weald.
D. aemula reaches the northern limit of its distribution in Britain.
20% of the world population of D. aemula is in the United Kingdom. [4]
A fern of shady, often rocky, woodlands, usually on acidic soils.
In Britain, it is generally found in NVC woodland community W17.
Matteuccia is a genus of ferns with one species: Matteuccia struthiopteris. The species epithet struthiopteris comes from Ancient Greek words στρουθίων (strouthíōn) "ostrich" and πτερίς (pterís) "fern".
Dryopteris, commonly called the wood ferns, male ferns, or buckler ferns, is a fern genus in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). There are about 300-400 species in the genus. The species are distributed in Asia, the Americas, Europe, Africa, and the Pacific islands, with the highest diversity in eastern Asia. It is placed in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Many of the species have stout, slowly creeping rootstocks that form a crown, with a vase-like ring of fronds. The sori are round, with a peltate indusium. The stipes have prominent scales.
Dryopteris expansa, the alpine buckler fern, northern buckler-fern or spreading wood fern, is a species of fern native to cool temperate and subarctic regions of the Northern Hemisphere, south at high altitudes in mountains to Spain and Greece in southern Europe, to Japan in eastern Asia, and to central California in North America. The species was first described from Germany. It prefers cool, moist mixed or evergreen forests and rock crevices on alpine slopes, often growing on rotting logs and tree stumps and rocky slopes. It is characteristically riparian in nature, and is especially associated with stream banks.
Ophioglossum azoricum, the small adder's-tongue fern or lesser adder's-tongue fern, is a small fern of the family Ophioglossaceae.
Dryopteris filix-mas, the male fern, is a common fern of the temperate Northern Hemisphere, native to much of Europe, Asia, and North America. It favours damp shaded areas in the understory of woodlands, but also shady places on hedge-banks, rocks, and screes. Near the northern limit of its distribution it prefers sunny, well-drained sites. It is much less abundant in North America than in Europe. The plant is sometimes referred to in ancient literature as worm fern, reflecting its former use against tapeworm.
Gymnocarpium dryopteris, the western oakfern, common oak fern, oak fern, or northern oak fern, is a deciduous fern of the family Cystopteridaceae. It is widespread across much of North America and Eurasia. It has been found in Canada, the United States, Greenland, China, Japan, Korea, Russia, and most of Europe.
Polystichum aculeatum, the hard shield-fern, is an evergreen fern native to Europe. It is most abundant in upland regions of the British Isles and western France, where it benefits from the combination of mild winters and moist summers, but also occurs more locally across most of Europe except northern Scandinavia, northern Russia; in the Mediterranean region it is confined to high altitudes. It grows on steep slopes in woodlands. it is sometimes considered an indicator of the presence of ancient woodlands.
Dryopteris erythrosora, the autumn fern or Japanese shield fern, is a species of fern in the family Dryopteridaceae, native to east Asia from China and Japan south to the Philippines, growing in light woodland shade on low mountains or hills.
Dryopteris affinis, the scaly male fern or golden-scaled male fern, is a fern native to western and southern Europe and southwestern Asia. It is most abundant on moist soils in woodlands in areas with high humidity, such as the British Isles and western France. In the Mediterranean region and the Caucasus it is confined to high altitudes.
Dryopteris carthusiana is a species of fern native to damp forests throughout the Holarctic Kingdom. It is known as the narrow buckler-fern in the United Kingdom, and as the spinulose woodfern in North America.
Dryopteris cristata is a species of fern native to wetlands throughout the Northern Hemisphere. It is known as crested wood fern, crested buckler-fern or crested shieldfern. This plant is a tetraploid species of hybrid origin, one parent being Dryopteris ludoviciana and the other being the unknown, apparently extinct species, dubbed Dryopteris semicristata, which is also one of the presumed parents of Dryopteris carthusiana. D. cristata in turn is one of the parents of Dryopteris clintoniana, another fern of hybrid origin.
Botrychium lunaria is a species of fern in the family Ophioglossaceae known by the common name moonwort or common moonwort. It is the most widely distributed moonwort, growing throughout the Northern Hemisphere across Eurasia and from Alaska to Greenland, as well as temperate parts of the Southern Hemisphere.
Amauropelta noveboracensis, the New York fern, is a perennial species of fern found throughout the eastern United States and Canada, from Louisiana to Newfoundland, but most concentrated within Appalachia and the Atlantic Northeast. New York ferns often forms spreading colonies within the forests they inhabit.
Polystichum lonchitis is a species of fern known by the common name northern hollyfern, or simply holly-fern. It is native to much of the Northern Hemisphere from Eurasia to Alaska to Greenland and south into mountainous central North America. It has stiff, glossy green, erect fronds and grows in moist, shady, rocky mountain habitats.
Cryptogramma crispa, the parsley fern, is an Arctic–alpine species of fern. It produces separate sterile and fertile fronds, up to 30 cm (12 in) tall, and is a pioneer species on acidic screes.
Gymnocarpium robertianum, the limestone fern or scented oakfern, is a fern of the family Cystopteridaceae.
Dryopteris dilatata, the broad buckler-fern, is a robust species of deciduous or semievergreen fern in the family Dryopteridaceae, native to Europe, particularly western and central Europe. In southern Europe, it is mostly found in mountainous regions. It is also found between the Black Sea and the Caspian Sea. It grows to 90 cm (35 in) tall by 120 cm (47 in) wide, with dark green tripinnate fronds, the ribs covered in brown scales.
Minwear Wood is a Site of Special Scientific Interest in Pembrokeshire, South Wales. It has been designated as a Site of Special Scientific Interest since January 1968 in an attempt to protect its fragile biological elements including ancient woodland of sessile oak with hazel and downy birch on the upper part of the Milford Haven estuary. The wood is structurally diverse and has scarce plants and epiphytic lichens. The site has an area of 13.69 hectares and is managed by Natural Resources Wales.
Hymenophyllum tunbrigense, the Tunbridge filmy fern or Tunbridge filmy-fern, is a small, fragile perennial leptosporangiate fern which forms large dense colonies of overlapping leaves from creeping rhizomes. The common name derives from the leaves which are very thin, only a single cell thick, and translucent, giving the appearance of a wet film. The evergreen fronds are bipinnatifid, deeply and irregularly dissected, about 3 to 6 cm long, 2 cm across with dark winged stipes. In contrast to the similar H. wilsonii the fronds are more divided, flattened, appressed to the substrate and tend to have a bluish tint.
Myriopteris aemula, the Texas lip fern or rival lip fern, is a moderately-sized fern of Texas and Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its leaves have a few hairs on upper and lower surfaces, or lack them entirely. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes as Cheilanthes aemula until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows on limestone rock.